Scarabaeidae

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File:Portrait of Beelte from Scarabaeidae family.jpg

On this high quality closeup, head anatomic details are well visible.

The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change. Several groups formerly treated as subfamilies have been elevated to family rank (e.g., Bolboceratidae, Geotrupidae, Glaresidae, Glaphyridae, Hybosoridae, Ochodaeidae, and Pleocomidae), and some reduced to lower ranks. The subfamilies listed in this article are in accordance with those in Catalog of Life (2023).<ref name=catlife/>

Description

File:Scarab Cartouche of Thutmosis III from Karnak.JPG

Sacred scarab in a cartouche of Thutmosis III from Karnak temple of Amun-Ra, Egypt

Scarabs are stout-bodied beetles, many with bright metallic colours, measuring between Template:Convert. They have distinctive, clubbed antennae composed of plates called lamellae that can be compressed into a ball or fanned out like leaves to sense odours. Many species are fossorial, with legs adapted for digging. In some groups males (and sometimes females) have prominent horns on the head and/or pronotum to fight over mates or resources.<ref name=bugguide /> The largest fossil scarabaeid was Oryctoantiquus borealis with a length of Template:Convert.<ref>Brett Ratcliffe, Dena M. Smith, Diane Marie Erwin. "Oryctoantiquus borealis, New Genus and Species from the Eocene of Oregon, U.S.A., the World's Oldest Fossil Dynastine and Largest Fossil Scarabaeid (Coleoptera: Scarabaeidae: Dynastinae)". 2009. The Coleopterists Bulletin 59(Mar 2005):127-135 DOI:10.1649/0010-065X(2005)059[0127:OBNGAS]2.0.CO;2</ref>

File:Curl grub.jpg

A scarab beetle grub from Australia.

The C-shaped larvae, called grubs, are pale yellow or white. Most adult beetles are nocturnal, although the flower chafers (Cetoniinae) and many leaf chafers (Rutelinae) are active during the day. The grubs mostly live underground or under debris, so are not exposed to sunlight. Many scarabs are scavengers that recycle dung, carrion, or decaying plant material.<ref>Template:Cite journal</ref> Others, such as the Japanese beetle, are plant-eaters, wreaking havoc on various crops and vegetation.

Some of the well-known beetles from the Scarabaeidae are Japanese beetles, dung beetles, June beetles, rose chafers (Australian, European, and North American), rhinoceros beetles, Hercules beetles and Goliath beetles.

Several members of this family have structurally coloured shells which act as left-handed circular polarisers; this was the first-discovered example of circular polarization in nature.<ref name="A.A. Michelson">Template:Cite journal</ref>

Ancient Egypt

In Ancient Egypt, the dung beetle now known as Scarabaeus sacer (formerly Ateuchus sacer) was revered as sacred.<ref>Template:Cite book</ref> Egyptian amulets representing the sacred scarab beetles were traded throughout the Mediterranean world.<ref name=bugguide>bugguide.net Family Scarabaeidae - Scarab Beetles</ref>