Editing Stork

{{Short description|Type of wading bird}}
{{Other uses}}
{{Automatic taxobox
| name = Stork
| fossil_range = [[Early Oligocene]] to present {{fossilrange|30|0}}
| image = Ringed white stork.jpg
| image_caption = [[White stork]] (''Ciconia ciconia'')
| display_parents = 2
| parent_authority = [[Charles Lucien Bonaparte|Bonaparte]], 1854<ref name="SN">{{cite web| url= http://taxonomicon.taxonomy.nl/TaxonTree.aspx?id=80129&tree=0.1| title=Systema Naturae 2000 / Classification, Class Aves | access-date=11 June 2012 | last=Brands | first=Sheila | date=14 August 2008 | work=Project: The Taxonomicon }}</ref>
| taxon = Ciconiidae
| authority = [[John Edward Gray|J. E. Gray]], 1840<ref name="SN" />
| subdivision_ranks = Genera
| subdivision =
*''[[Anastomus]]''
*''[[Ciconia]]''
*''[[Ephippiorhynchus]]''
*''[[Jabiru]]''
*''[[Leptoptilos]]''
*''[[Mycteria]]''
| synonyms =
}}

'''Storks''' are large, long-legged, long-necked [[wading bird]]s with long, stout [[Beak|bills]]. They belong to the [[family (biology)|family]] '''Ciconiidae''', and make up the order '''Ciconiiformes''' {{IPAc-en|s|ᵻ|ˈ|k|oʊ|n|i|.|ᵻ|f|ɔr|m|iː|z}}. Ciconiiformes previously included a number of other families, such as [[heron]]s and [[ibis]]es, but those families have been moved to other orders.<ref>{{cite web|url=https://www.gbif.org/species/100220333|title=Anastomus lamelligerus subsp. lamelligerus|website=www.gbif.org|access-date=2018-03-22|archive-date=2018-08-03|archive-url=https://web.archive.org/web/20180803194327/https://www.gbif.org/species/100220333|url-status=dead}}</ref> 

Storks dwell in many regions and tend to live in drier habitats than the closely related [[heron]]s, [[spoonbill]]s and [[ibis]]es; they also lack the [[powder down]] that those groups use to clean off [[fish]] slime. Bill-clattering is an important mode of [[Animal communication|communication]] at the nest. Many species are [[bird migration|migratory]]. Most storks eat [[frog]]s, [[fish]], [[insect]]s, [[earthworm]]s, small [[bird]]s and small [[mammal]]s. There are 20 living [[species]] of storks in six [[genera]].

Various terms are used to refer to groups of storks,<ref>[http://oilspillwildlife.org/index.php?option=com_content&view=article&id=70 ''About the Wood Stork: Denizens of the Wetlands''] {{Webarchive|url=https://web.archive.org/web/20110727141943/http://oilspillwildlife.org/index.php?option=com_content&view=article&id=70 |date=2011-07-27 }}, Accessed on 13.12.2010</ref> two frequently used ones being a ''muster'' of storks and a ''phalanx'' of storks.

Storks tend to use [[Lift (soaring)|soaring]], gliding flight, which conserves energy.  Soaring requires [[thermal]] air currents. [[Ottomar Anschütz]]'s famous 1884 album of photographs of storks inspired the design of [[Otto Lilienthal]]'s experimental [[Glider aircraft|gliders]] of the late nineteenth century. Storks are heavy, with wide [[wingspan]]s: the [[marabou stork]], with a wingspan of {{cvt|3.2|m|ftin}} and weight up to {{cvt|8|kg|lb|0}}, joins the [[Andean condor]] in having the widest wingspan of all living land birds.

Their [[bird nest|nests]] are often very large and may be used for many years. Some nests have been known to grow to over {{convert|2|m}} in diameter and about {{convert|3|m}} in depth. All storks were once thought to be [[monogamy in animals|monogamous]], but this is only partially true. While storks are generally socially monogamous, some species exhibit regular extra-pair breeding.<ref>{{Cite journal |last1=Turjeman |first1=Sondra F. |last2=Centeno-Cuadros |first2=Alejandro |last3=Eggers |first3=Ute |last4=Rotics |first4=Shay |last5=Blas |first5=Julio |last6=Fielder |first6=Wolfgang |last7=Kaatz |first7=Michael |last8=Jeltsch |first8=Florian |last9=Wikelski |first9=Martin |last10=Ran |first10=Nathan |date=2016 |title=Extra-pair paternity in the socially monogamous white stork (Ciconia ciconia) is fairly common and independent of local density |journal=Scientific Reports |volume=6 |pages=27976 |doi=10.1038/srep27976|pmid=27328982 |pmc=4916429 |bibcode=2016NatSR...627976T }}</ref>

Popular conceptions of storks' fidelity, [[serial monogamy]], and doting parental care contribute to their prominence in [[mythology]] and culture, especially in western folklore as the deliverers of newborn humans.<ref>{{Cite journal |last1=Margolis |first1=Marvin |last2=Parker |first2=Philip |date=1972 |title=The stork fable - some psychodynamic considerations |journal=Journal of the American Psychoanalytic Association |volume=20 |issue=3 |pages=494–511 |doi=10.1177/000306517202000304|pmid=4116100 }}</ref>

All 20 stork species have been assessed by the IUCN and carry a confident Red List status. However, the assessment for several species were based on incorrect assumptions and a general absence of sound information on stork habits.<ref>{{Cite journal |last1=Gula |first1=Jonah |last2=Sundar |first2=K. S. Gopi |last3=Willows-Munro |first3=Sandi |last4=Downs |first4=Colleen T. |date=2023 |title=The state of stork research globally: A systematic review |journal=Biological Conservation |volume=280 |issue=April 2023 |pages=109969|doi=10.1016/j.biocon.2023.109969 |bibcode=2023BCons.28009969G |doi-access=free }}</ref>

==Etymology==
The word "stork" was first used in its current sense by at least the 12th century in [[Middle English]].<ref name=":5">{{Cite Merriam-Webster|stork|accessdate=2023-07-29}}</ref> It is derived from the [[Old English]] word "storc", which itself comes from the hypothesised [[Proto-Germanic]] {{Langx|gem-x-proto|*stork|links=|label=none}} and ultimately the [[Proto-Indo-European]] ''{{PIE|*sr̥ǵos}}''.<ref name=":5" /><ref>{{Cite journal |last=Blažek |first=Václav |date=2003 |title=A Tocharian key to the etymology of the bird-name *sr̥ǵos-"stork" |url=https://books.google.com/books?id=RL7fDwAAQBAJ&dq=stork+etymology&pg=PA63 |journal=[[Tocharian and Indo-European Studies]] |volume=10 |pages=15–16|isbn=9788021076457 }}</ref> The name refers to the rigid posture of storks, a meaning reflected in the related word [[wiktionary:stark|stark]], which is derived from the Old English "stearc".<ref name="OEDstork">{{Cite OED|term=stork|id=3439296373|access-date=2023-07-29}}</ref> Several species of storks are known by other common names. The [[jabiru]] is named after the [[Tupi–Guarani languages|Tupí-Guarani]] words meaning "that which has" and "swollen", referring to its thickset neck.<ref name="OEDjabiru">{{Cite OED|term=jabiru|id=1060893535|access-date=2023-07-29}}</ref> The [[marabou stork]] is named after the Arabic word for holy man, ''murābiṭ'', due to the perceived holy nature of the species.<ref name="OEDmarabou">{{Cite OED|term=marabou stork|id=5740365926|access-date=2023-07-29}}</ref> The [[Leptoptilos|adjutants]] are named after [[Adjutant|the military rank]], referring to their stiff, military-like gait.<ref name="OEDadjutant">{{Cite OED|term=adjutant stork|id=9863034263|access-date=2023-07-29}}</ref>

==Systematics==
{{See also|List of storks}}
A DNA study found that the families [[Ardeidae]], [[Balaenicipitidae]], [[Scopidae]] and the [[Threskiornithidae]] belong to the [[Pelecaniformes]]. This would make [[Ciconiidae]] the only group.<ref>{{Cite journal | doi=10.1016/j.ympev.2013.03.021| pmid=23562800| title=Beyond phylogeny: Pelecaniform and ciconiiform birds, and long-term niche stability| journal=Molecular Phylogenetics and Evolution| volume=68| issue=2| pages=229–238| year=2013| last1=Gibb| first1=Gillian C.| last2=Kennedy| first2=Martyn| last3=Penny| first3=David| bibcode=2013MolPE..68..229G}}</ref><ref>{{cite journal|doi=10.1093/gbe/evv213|pmid=26527652|title=Determining the Position of Storks on the Phylogenetic Tree of Waterbirds by Retroposon Insertion Analysis|journal=Genome Biology and Evolution|volume=7|issue=12|pages=3180–3189|year=2015|last1=Kuramoto|first1=Tae|last2=Nishihara|first2=Hidenori|last3=Watanabe|first3=Maiko|last4=Okada|first4=Norihiro|pmc=4700946}}</ref>

Storks were distinct and possibly widespread by the [[Oligocene]]. Like most families of aquatic birds, storks seem to have arisen in the [[Paleogene|Palaeogene]], maybe 40–50 [[million years ago]] (mya). For the fossil record of living genera, documented since the [[Middle Miocene]] (about 15 mya) at least in some cases, see the genus articles.

No species or subspecies of stork is known to have gone [[extinct]] in historic times. A systematic literature review uncovered nearly 1,000 papers on storks, but showed most stork species to lack scientific understanding suggesting that many species should be classified as Data Deficient on the IUCN Red List. A ''Ciconia'' bone found in a [[rock shelter]] on the island of [[Réunion]] was probably of a bird taken there as food by early settlers; no known account mentions the presence of storks on the [[Mascarene Islands]].

=== Phylogeny ===
The following phylogeny is recognized by the International Ornithological Congress, partially based on de Sousa ''et al'' (2023):<ref>{{Cite journal |last1=de Sousa |first1=Rodrigo Petry Corrêa |last2=Campos |first2=Paula Sabrina Bronze |last3=dos Santos |first3=Michelly da Silva |last4=O’Brien |first4=Patricia Caroline |last5=Ferguson-Smith |first5=Malcolm Andrew |last6=de Oliveira |first6=Edivaldo Herculano Corrêa |date=April 2023 |title=Cytotaxonomy and Molecular Analyses of Mycteria americana (Ciconiidae: Ciconiiformes): Insights on Stork Phylogeny |journal=Genes |language=en |volume=14 |issue=4 |pages=816 |doi=10.3390/genes14040816 |issn=2073-4425 |pmc=10138051 |pmid=37107574 |doi-access=free }}</ref><ref>{{Cite web |title=Storks, frigatebirds, boobies, darters, cormorants – IOC World Bird List |url=https://www.worldbirdnames.org/new/bow/storks/ |access-date=2023-07-30 |website=www.worldbirdnames.org}}</ref>
{{Clade|style=font-size:100%
   |1={{clade
        |1=[[African openbill]] (''Anastomus lamelligerus'')
        |2=[[Asian openbill]] ''(Anastomus oscitans'')
  }}
   |2={{clade
        |1={{clade
               |1=[[Marabou stork]] (''Leptoptilos crumenifer'')
              |2={{clade
                   |1=[[Lesser adjutant]] (''Leptoptilos javanicus'')
                   |2=[[Greater adjutant]] (''Leptoptilos dubius'')
       }}
}}
        |2={{clade
        |1={{clade
              |1=[[Wood stork]] (''Mycteria americana'')
              |2={{clade
                   |1=[[Yellow-billed stork]] (''Mycteria ibis'')
                   |2={{clade
                        |1=[[Painted stork]] (''Mycteria leucocephala'')
                       |2=[[Milky stork]]
(''Mycteria cinerea'')
        }}
}}
}}
        |2={{clade
            |1={{clade
                 |1=[[Jabiru]] (''Jabiru mycteria'')
                 |2={{clade
                      |1=[[Saddle-billed stork]] (''Ephippiorhynchus senegalensis'')
                     |2=[[Black-necked stork]] (''Ephippiorhynchus asiaticus'')
            }}
       }}
            |2={{clade
                  |1={{clade
                      |1=[[Abdim's stork]] (''Ciconia abdimii'')
                     |2={{clade
                         |1=[[Asian woolly-necked stork]] (''Ciconia episcopus'')
                        |2=[[Storm's stork]] (''Ciconia stormi'')
                     }}
                }}
                |2={{clade
                   |1=[[Black stork]] (''Ciconia   nigra'')
                   |2={{clade
                         |1=[[Maguari stork]] (''Ciconia maguari'')
                         |2={{clade
                               |1=[[African woolly-necked stork]] (''Ciconia microscelis'')
                               |2={{clade
                                   |1=[[White stork]] (''Ciconia ciconia'')
                                   |2=[[Oriental stork]] (''Ciconia boyciana'')
                               }}
                           }}
       }}
}}
}}
}}
}}
}}
}}

===Fossil storks===
*Genus ''[[Palaeoephippiorhynchus]]'' ([[fossil]]: Early Oligocene of [[Fayyum]], Egypt)<!-- ActaPaleontolPol50:549 -->
*Genus ''[[Grallavis]]'' ([[fossil]]: Early Miocene of Saint-Gérand-le-Puy, France, and Djebel Zelten, Libya) – may be same as ''[[Prociconia]]''
*Ciconiidae gen. et sp. indet. (Ituzaingó Late Miocene of Paraná, Argentina)<ref group=note>[[Tarsometatarsus]] fragments somewhat similar to ''Mycteria''</ref><ref name=Cione>{{cite book|last1=Cione|first1= Alberto Luis|last2= de las Mercedes Azpelicueta|first2= María|last3= Bond|first3= Mariano|last4= Carlini|first4= Alfredo A.|last5= Casciotta|first5= Jorge R.|last6= Cozzuol|first6= Mario Alberto|last7= de la Fuente|first7= Marcelo|last8= Gasparini|first8= Zulma|last9= Goin|first9= Francisco J.|last10= Noriega|first10= Jorge|last11= Scillatoyané|first11= Gustavo J.|last12= Soibelzon | first12 = Leopoldo|last13= Tonni|first13= Eduardo Pedro|last14= Verzi|first14= Diego |last15= Guiomar Vucetich|first15= María|date=2000|chapter= Miocene vertebrates from Entre Ríos province, eastern Argentina|language =en, es|editor1-last= Aceñolaza|editor1-first= F.G. |editor2-last=  Herbst|editor2-first= R.|title= El Neógeno de Argentina|publisher=INSUGEO|series= Serie Correlación Geológica|volume =14|pages = 191–237|chapter-url =  http://www.unt.edu.ar/fcsnat/INSUGEO/libro_neogeno/neogeno_argentina.pdf |url-status = dead|archive-url=https://web.archive.org/web/20080228004955/http://www.unt.edu.ar/fcsnat/INSUGEO/libro_neogeno/neogeno_argentina.pdf |archive-date=2008-02-28 }}</ref><ref name=Noriega>Noriega, Jorge Ignacio & Cladera, Gerardo (2005). ''First Record of Leptoptilini (Ciconiiformes: Ciconiidae) in the Neogene of South America''. Abstracts of Sixth International Meeting of the Society of Avian Paleontology and Evolution: 47. [http://nrm.museum/ve/birds/sape/SAPE_abstracts_2004.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20080228004954/http://nrm.museum/ve/birds/sape/SAPE_abstracts_2004.pdf |date=2008-02-28 }}</ref>
*Ciconiidae gen. et sp. indet. (Puerto Madryn Late Miocene of Punta Buenos Aires, Argentina)<ref group=note>Specimen [[Museo Paleontológico Egidio Feruglio|MEF]] 1363: Incomplete skeleton of a large stork somewhat similar to ''Jabiru'' but apparently more [[plesiomorph]]ic</ref><ref name= Noriega/>
*Genus ''[[Prociconia]]'' ([[fossil]]: Late Pleistocene of Brazil) – may belong to modern genus ''Jabiru'' or ''Ciconia''<!-- Evolution39:1174 -->
*Genus ''[[Pelargosteon]]'' ([[fossil]]: Early Pleistocene of Romania)
*Ciconiidae gen. et sp. indet. – formerly ''Aquilavus/Cygnus bilinicus'' ([[fossil]]: Early Miocene of Břešťany, Czech Republic)
*cf. ''[[Leptoptilos]]'' gen. et sp. indet. – formerly ''L. siwalicensis'' ([[fossil]]: Late Miocene? – Late Pliocene of Siwalik, India)<ref name="Louchart">Specimens [[British Museum of Natural History|BMNH]] 39741 (holotype, left [[Anatomical terms of location#Proximal and distal|proximal]] [[tarsometatarsus]]) and BMNH 39734 (right [[Anatomical terms of location#Proximal and distal|distal]] [[tibiotarsus]]). Similar to ''Ephippiorhynchus'' and ''Leptotilos'', may be from a small female of ''[[Leptotilos falconeri]]'', from ''L. dubius'', or from another species: {{cite journal |author1=Louchart, Antoine |author2=Vignaud, Patrick |author3=Likius, Andossa |author4=Brunet, Michel |author5=White, Tim D. |name-list-style=amp |year=2005 |title=A large extinct marabou stork in African Pliocene hominid sites, and a review of the fossil species of ''Leptoptilos'' |url=http://www.app.pan.pl/acta50/app50-549.pdf |url-status=dead |journal=Acta Palaeontologica Polonica |volume=50 |issue=3 |pages=549–563 |archive-url=https://web.archive.org/web/20080228004954/http://www.app.pan.pl/acta50/app50-549.pdf |archive-date=2008-02-28 |access-date=2008-01-15}}</ref>
*Ciconiidae gen. et sp. indet. ([[fossil]]: Late Pleistocene of San Josecito Cavern, Mexico)<ref name=Steadman>[[Anatomical terms of location#Proximal and distal|Distal]] [[radius (bone)|radius]] of a mid-sized ''Ciconia'' or smallish ''Mycteria'': {{cite journal|author1=Steadman, David W. |author2=Arroyo-Cabrales, Joaquin |author3=Johnson, Eileen |author4=Guzman, A. Fabiola  |name-list-style=amp |year=1994|title= New Information on the Late Pleistocene Birds from San Josecito Cave, Nuevo León, Mexico|journal=[[Condor (journal)|Condor]]|volume=96|issue=3|pages= 577–589|url=http://sora.unm.edu/sites/default/files/journals/condor/v096n03/p0577-p0589.pdf|doi=10.2307/1369460|jstor=1369460 }}</ref>
*''Ciconia nana'' ([[fossil]]: Pleistocene of Darling Downs, Queensland, Australia, and ''Ciconia louisebolesae'' [[fossil]]: Olig-Miocene of Riversleigh WHA, Queensland, Australia<ref>Boles. [[A review of the Australian fossil storks of the genus ''Ciconia'' (Aves:Ciconiidae), with a description of a new species]]|journal=[[Records of the Australian Museum]] (2005) Vol=57, pages= 165–178</ref>

The fossil genera ''Eociconia'' (Middle Eocene of China) and ''Ciconiopsis'' (Deseado Early Oligocene of Patagonia, Argentina) are often tentatively placed with this family. A "ciconiiform" fossil fragment from the Touro Passo Formation found at Arroio Touro Passo ([[Rio Grande do Sul]], [[Brazil]]) might be of the living [[wood stork]] ''M. americana''; it is at most of [[Late Pleistocene]] age, a few 10,000s of years.<ref name=Schmaltz>Schmaltz Hsou, Annie (2007). ''O estado atual do registro fóssil de répteis e aves no Pleistoceno do Estado do Rio Grande do Sul, Brasil'' ["The current state of the fossil record of Pleistocene reptiles and birds of Rio Grande do Sul"]. Talk held on 2007-JUN-20 at ''Quaternário do RS: integrando conhecimento'', Canoas, Rio Grande do Sul, Brazil. [http://www6.ufrgs.br/alpp/Resumos_Quaternario_RS.pdf PDF abstract] {{Webarchive|url=https://web.archive.org/web/20170708084133/http://www.ufrgs.br/alpp/Resumos_Quaternario_RS.pdf |date=2017-07-08 }}</ref>

[[File:Dvorac Bisag - roda.jpg|thumb|European white stork in a nest in Bisag, [[Croatia]]]]

==Morphology==
[[File:Yellow-billed Stork (Mycteria ibis) (12011503884).jpg|thumb|''Mycteria'' storks, like this yellow-billed stork, have sensitive bills that allow them to hunt by touch]]
Storks range in size from the marabou, which stands {{convert|152|cm|in|abbr=on}} tall and can weigh {{convert|8.9|kg|lb|frac=4|abbr=on}}, to the Abdim's stork, which is only {{convert|75|cm|in|-1|abbr=on}} high and weighs only {{convert|1.3|kg|lb|frac=4|abbr=on}}. Their shape is superficially similar to the herons, with long legs and necks, but they are more heavy-set. There is some [[sexual dimorphism]] (differences between males and females) in size, with males being up to 15% bigger than females in some species (for example the saddle-billed stork), but almost no difference in appearance. The only difference is in the colour of the [[Iris (anatomy)|iris]] of the two species in the genus ''[[Ephippiorhynchus]]''.<ref name="HBW" />

The [[beak|bills]] of storks are large to very large, and vary considerably between the genera. The shape of the bills is linked to the diet of the different species. The large bills of the ''Ciconia'' storks are the least specialized. Larger are the massive and slightly upturned bills of the ''Ephippiorhynchus'' and the jabiru. These have evolved to hunt for fish in shallow water. Larger still are the massive daggers of the two adjutants and marabou (''Leptoptilos''), which are used to feed on carrion and in defense against other scavengers, as well as for taking other prey.<ref name="HBW" /> The long, ibis-like downcurved bills of the ''Mycteria'' storks have sensitive tips that allow them to detect prey by touch (tactilocation) where cloudy conditions would not allow them to see it.<ref>{{cite journal|last1=Coulter|first1=Malcolm C.|last2=Bryan|first2=A. Lawrence|title=Foraging Ecology of Wood Storks (Mycteria americana) in East-Central Georgia I. Characteristics of Foraging Sites|journal=Colonial Waterbirds|date=1 January 1993|volume=16|issue=1|pages=59–70|doi=10.2307/1521557|jstor=1521557}}</ref> The most specialised bills of any storks are those of the two openbills (''Anastomus''), which as their name suggests, is open in the middle when their bill is closed. These bills have evolved to help openbills feed on their primary prey item, aquatic snails.<ref name="openbill diet">{{cite journal|last1=Kahl|first1=M. P.|title=Food and feeding behavior of Openbill Storks|journal=Journal of Ornithology|date=January 1971|volume=112|issue=1|pages=21–35|doi=10.1007/BF01644077|bibcode=1971JOrni.112...21K |s2cid=1484358}}</ref>

Although it is sometimes reported that storks lack [[Syrinx (bird anatomy)|syrinxes]] and are mute,<ref>{{cite web|last1=Spring Alive|title=A bird without voice|url=http://www.springalive.net/world/spring_news/Birdwithoutvoice|access-date=7 March 2017}}</ref> they do have syrinxes,<ref name ="Griffiths">{{cite journal|last1=Griffiths|first1=Carole S.|title=Monophyly of the Falconiformes Based on Syringeal Morphology|journal=The Auk|date=October 1994|volume=111|issue=4|pages=787–805|doi=10.2307/4088811|jstor=4088811|s2cid=53388019|doi-access=free}}</ref> and are capable of making some sounds, although they do not do so often.<ref name="HBW" /><ref>{{Cite journal|last=Sundar|first=K.S. Gopi|date=2005|title=An instance of mortality and notes on behaviour of black-necked storks Ephippiorhynchus asiaticus|journal=Journal of the Bombay Natural History Society|volume=102|issue=1|pages=99–102}}</ref>  The syrinxes of storks are "variably degenerate" however,<ref name ="Griffiths"/> and the syringeal membranes of some species are found between tracheal rings or cartilage, an unusual arrangement shared with the [[Ovenbird (family)|ovenbirds]].<ref>{{cite journal|last1=Casey|first1=Richard M.|last2=Gaunt|first2=Abbot S.|title=Theoretical models of the avian syrinx|journal=Journal of Theoretical Biology|date=September 1985|volume=116|issue=1|pages=45–64|doi=10.1016/S0022-5193(85)80130-2|bibcode=1985JThBi.116...45C}}</ref>

==Distribution and habitat==
[[File:Lesser Adjutant (Leptoptilos javanicus) - Flickr - Lip Kee (4).jpg|thumb|[[Lesser adjutant]]s will forage in marine habitats, unlike most storks]]
[[File:Marabou stork at Etosha National Park, Namibia.jpg|thumb|[[Marabou stork|Marabou]] stork at [[Etosha National Park]] in [[Namibia]]]]
{{See also|List of Ciconiiformes by population}}
Storks have a nearly [[cosmopolitan distribution]], being absent from the poles, most of North America and large parts of Australia. The centres of stork diversity are in tropical Asia and sub-Saharan Africa, with eight and six breeding species respectively. Just three species are present in the New World: [[wood stork]], [[maguari stork]] and [[jabiru]], which is the tallest flying bird of the Americas. Two species, [[white stork|white]] and [[black stork]], reach Europe and western temperate Asia, while one species, [[Oriental stork]], reaches temperate areas of eastern Asia, and one species, [[black-necked stork]], is found in Australasia.<ref name = "HBW">del Hoyo, J. Elliott, A. & Sargatal, J. (1992). ''Handbook of the Birds of the World. Volume 1: Ostrich to Ducks. Lynx Edicions''. {{ISBN|84-87334-10-5}}.</ref>

Storks are more diverse and common in the tropics, and the species that live in temperate climates for the most part [[bird migration|migrate]] to avoid the worst of winter. They are fairly diverse in their habitat requirements. Some species, particularly the ''[[Mycteria ]]'' "wood storks" and ''[[Anastomus]]'' openbills, are highly dependent on water and aquatic prey, but many other species are far less dependent on this habitat type, although they will frequently make use of it. Species like the [[Marabou stork|marabou]] and [[Abdim's stork]] will frequently be found foraging in open grasslands of savannah. Preferred habitats include flooded grasslands, light woodland, marshes and paddyfields, [[wet meadow]]s, river backwaters and ponds. Many species will select shallow pools, particularly when lakes or rivers are drying out, as they concentrate prey and make it harder for prey to escape, or when monsoonal rainfall increases water depth of larger waterbodies.<ref name="HBW" /><ref name=":0">{{Cite journal|last=Sundar|first=K.S. Gopi|date=2006|title=Flock Size, Density and Habitat Selection of Four Large Waterbirds Species in an Agricultural Landscape in Uttar Pradesh, India: Implications for Management|url=https://bioone.org/journals/waterbirds/volume-29/issue-3/1524-4695(2006)29%5b365%3aFSDAHS%5d2.0.CO%3b2/Flock-Size-Density-and-Habitat-Selection-of-Four-Large-Waterbirds/10.1675/1524-4695(2006)29%5b365:FSDAHS%5d2.0.CO;2.short|journal=Waterbirds|volume=29|issue=3|pages=365–374|doi=10.1675/1524-4695(2006)29[365:FSDAHS]2.0.CO;2|s2cid=198154724 }}</ref> Some species like the woolly-necked storks and lesser adjutant storks have adapted to changing crops of tropical agricultural landscapes that enables them to remain resident despite the transformations brought about by seasonal crops.<ref name=":4">{{Cite journal |last1=Katuwal |first1=Hem B. |last2=Sundar |first2=K. S. Gopi |last3=Zhang |first3=M |last4=Rimal |first4=B |last5=Baral |first5=H. S. |last6=Sharma |first6=H. P. |last7=Ghimire |first7=P |last8=Hughes |first8=A. C. |last9=Quan |first9=R.-C. |date=2022 |title=Factors affecting the breeding ecology of the globally threatened Lesser Adjutant (Leptoptilos javanicus) in agricultural landscapes of Nepal |url=https://ace-eco.org/vol17/iss2/art15/ |journal=Avian Conservation and Ecology |volume=17 |issue=2 |pages=15|doi=10.5751/ACE-02235-170215 |s2cid=251862746 |doi-access=free }}</ref><ref name=":6">{{Cite journal |last1=Kittur |first1=Swati |last2=Sundar |first2=K. S. Gopi |date=2021 |title=Of irrigation canals and multifunctional agroforestry: Traditional agriculture facilitates Woolly-necked Stork breeding in a north Indian agricultural landscape |journal=Global Ecology and Conservation |volume=30 |issue=October 2021 |pages=e01793|doi=10.1016/j.gecco.2021.e01793 |s2cid=239153561 |doi-access=free |bibcode=2021GEcoC..3001793K }}</ref><ref>{{Cite journal |last1=Ghimire |first1=P |last2=Pandey |first2=N |last3=Timilsina |first3=Y. P. |last4=Bist |first4=B. S. |last5=Sundar |first5=K. S. Gopi |date=2021 |title=Woolly-Necked Stork (Ciconia episcopus) activity budget in lowland Nepal's farmlands: the influence of wetlands, seasonal crops, and human proximity |url=https://bioone.org/journals/waterbirds/volume-44/issue-4/063.044.0403/Woolly-Necked-Stork-Ciconia-episcopus-Activity-Budget-in-Lowland-Nepals/10.1675/063.044.0403.short |journal=Waterbirds |volume=44 |issue=4 |pages=415–424|doi=10.1675/063.044.0403 |s2cid=251161403 }}</ref><ref>{{Cite journal |last1=Sundar |first1=K. S. Gopi |last2=Maharjan |first2=B |last3=Koju |first3=Roshila |last4=Kittur |first4=Swati |last5=Gosai |first5=Kamal R. |date=2016 |title=Factors affecting provisioning times of two stork species in lowland Nepal |journal=Waterbirds |volume=39 |issue=4 |pages=365–374|doi=10.1675/063.039.0406 |s2cid=90680383 |doi-access=free }}</ref> In South Africa, the woolly-necked storks have adapted to artificial feeding and now largely nest on trees in gardens with swimming pools.<ref name=":7">{{Cite journal |last1=Thabethe |first1=V |last2=McPherson |first2=S |last3=Downs |first3=C. T. |date=2021 |title=Diet of nestling African woolly-necked storks in suburban areas of KwaZulu-Natal, South Africa |url=https://onlinelibrary.wiley.com/doi/10.1111/aje.12859 |journal=African Journal of Ecology |volume=59 |issue=3 |pages=769–772|doi=10.1111/aje.12859 |bibcode=2021AfJEc..59..769T |s2cid=233950996 }}</ref>

Less typical habitats include the dense temperate forests used by European [[Black stork|black storks]], or the rainforest habitat sought by [[Storm's stork]] in South East Asia. They generally avoid marine habitats, with the exception of the [[lesser adjutant]], [[milky stork]] and [[wood stork]], all of which forage in mangroves, lagoons and estuarine mudflats. A  number of species, especially woolly-necked storks, black-necked storks, Asian openbills and lesser adjutant Storks in south Asia, have adapted to highly modified human habitats, for foraging and breeding.<ref name=":0" /><ref name=":1">{{Cite journal|last1=Koju|first1=Roshila|last2=Maharjan|first2=Bijay|last3=Gosai|first3=Kamal Raj|last4=Kittur|first4=Swati|last5=Sundar|first5=K.S. Gopi|date=2019|title=Ciconiiformes nesting on trees in cereal-dominated farmlands: importance of scattered trees for heronries in lowland Nepal|journal=Waterbirds|volume=42|issue=4|pages=355–365|doi=10.1675/063.042.0401|s2cid=210861485|doi-access=free}}</ref><ref name=":2">{{Cite journal|last1=Sundar|first1=K.S. Gopi|last2=Maharjan|first2=Bijay|last3=Koju|first3=Roshila|last4=Kittur|first4=Swati|last5=Gosai|first5=Kamal Raj|date=2016|title=Factors affecting provisioning times of two stork species in lowland Nepal|journal=Waterbirds|volume=39|issue=4|pages=365–374|doi=10.1675/063.039.0406|s2cid=90680383|doi-access=free}}</ref><ref name=":3">{{Cite journal|last1=Kittur|first1=Swati|last2=Sundar|first2=K. S. Gopi|date=2021|title=Of irrigation canals and multifunctional agroforestry: Traditional agriculture facilitates Woolly-necked Stork breeding in a north Indian agricultural landscape|journal=Global Ecology and Conservation|volume=30|pages=e01793|doi=10.1016/j.gecco.2021.e01793| issn=2351-9894|s2cid=239153561|doi-access=free|bibcode=2021GEcoC..3001793K }}</ref> In the absence of persecution several stork species breed close to people, and species such as the marabou, greater adjutant, and white stork feed at landfill sites.<ref name="HBW" /><ref>{{cite journal|last1=Tortosa|first1=F. S.|last2=Caballero|first2=J. M.|last3=Reyes-López|first3=J.|title=Effect of Rubbish Dumps on Breeding Success in the White Stork in Southern Spain|journal=Waterbirds|date=March 2002|volume=25|issue=1|pages=39–43|doi=10.1675/1524-4695(2002)025[0039:EORDOB]2.0.CO;2|s2cid=85703087 }}</ref>

===Migration and movements===
[[File:Abdim's Stork (Ciconia abdimii) (7011390701).jpg|thumb|Abdim's storks are regular intra-African migrants]]
Storks vary in their tendency towards migration. Temperate species like the white stork, black stork and Oriental stork undertake long annual migrations in the winter. The routes taken by these species have developed to avoid long distance travel across water, and from Europe this usually means flying across the Straits of Gibraltar or east across the Bosphorus and through Israel and the Sinai.<ref name="HBW" /> Studies of young birds denied the chance to travel with others of their species have shown that these routes are at least partially learnt, rather than being innate as they are in [[passerine]] migrants.<ref name="migration">{{cite journal|last1=Chernetsov|first1=N.|last2=Berthold|first2=P.|last3=Querner|first3=U.|title=Migratory orientation of first-year white storks (Ciconia ciconia): inherited information and social interactions|journal=Journal of Experimental Biology|date=22 February 2004|volume=207|issue=6|pages=937–943|doi=10.1242/jeb.00853|doi-access=free|pmid=14766952}}</ref> Migrating black storks are split between those that make stopovers on the migration between Europe and their wintering grounds in Africa, and those that do not.<ref>{{cite journal|last1=Chevallier|first1=D.|last2=Le Maho|first2=Y.|last3=Brossault|first3=P.|last4=Baillon|first4=F.|last5=Massemin|first5=S.|title=The use of stopover sites by Black Storks (Ciconia nigra) migrating between West Europe and West Africa as revealed by satellite telemetry|journal=Journal of Ornithology|date=5 June 2010|volume=152|issue=1|pages=1–13|doi=10.1007/s10336-010-0536-6|s2cid=21513063}}</ref>

The Abdim's stork is another migrant, albeit one that migrates within the tropics. It breeds in northern Africa, from Senegal to the Red Sea, during the wet season, and then migrates to Southern Africa.<ref>{{cite journal|last1=Adjakpa|first1=Jacques Boco|title=The breeding biology of Abdim's Stork in the far north of Benin|journal=Ostrich|date=January 2000|volume=71|issue=1–2|pages=61–63|doi=10.1080/00306525.2000.9639869|s2cid=84995962}}</ref> Many species that are not regular migrants will still make smaller movements if circumstances require it; others may migrate over part of their range. This can also include regular commutes from nesting sites to feeding areas. Wood storks have been observed feeding {{convert|130|km|mi|-1|abbr=on}} from their breeding colony.<ref name="HBW" />

==Behaviour==
===Feeding and diet===
[[File:African openbill, Anastomus lamelligerus, Chobe National Park, Botswana (32390774275).jpg|thumb|African openbill foraging in shallow water]]
Storks are carnivorous predators, taking a range of reptiles, small mammals, insects, fish, amphibians and other small invertebrates. Storks usually hunt for animals in shallow water. Any plant material consumed is usually by accident. ''Mycteria'' storks are specialists in feeding on aquatic vertebrates, particularly when prey is concentrated by lowering water levels or flooding into shallows. On marine mudflats and mangrove swamps in [[Sumatra]], milky storks feed on [[mudskipper]]s, probing the burrow with the bill and even the whole head into the mud.{{Citation needed|date=April 2021}} The characteristic feeding method involves standing or walking in shallow water and holding the bill submerged in the water. When contact is made with prey the bill reflexively snaps shut in 25 milliseconds, one of the fastest reactions known in any vertebrate. The reaction is able to distinguish between prey items and inanimate objects like branches, although the exact mechanism is unknown.<ref>{{cite journal|last1=Kahl|first1=M. P.|last2=Peacock|first2=L. J.|title=The Bill-snap Reflex : a Feeding Mechanism in the American Wood Stork|journal=Nature|date=3 August 1963|volume=199|issue=4892|pages=505–506|doi=10.1038/199505a0|pmid=14058622|bibcode=1963Natur.199..505K|s2cid=332546}}</ref><ref name="HBW" />

Openbills are specialists in freshwater molluscs, particularly apple snails. They feed in small groups,<ref name=":0" /> and sometimes African openbills ride on the backs of hippos while foraging. Having caught a snail it will return to land or at least to the shallows to eat it. The fine tip of the bill of the openbills is used to open the snail, and the saliva has a narcotic effect, which causes the snail to relax and simplifies the process of extraction.<ref name="HBW" />

The other genera of storks are more generalised. ''Ciconia'' storks are very generalised in their diets, and some species including Abdim's stork and marabous will feed in large flocks on swarms of locusts and at wildfires.<ref name="HBW" /><ref>{{cite journal|last1=Dean|first1=G. J. W.|title=Stork and egret as predators of the red locust in the Rukwa Valley outbreak area|journal=Ostrich|date=June 1964|volume=35|issue=2|pages=95–100|doi=10.1080/00306525.1964.9633490|bibcode=1964Ostri..35...95D }}</ref> This is why white storks and Abdim's storks are known as "grasshopper birds". ''Ephippiorhynchus'' are carnivorous though have a very diverse diet when living on human modified habitats such as agricultural landscapes.<ref>{{Cite journal|last=Sundar|first=K.S. Gopi|date=2011|title=Farmland foods: Black-necked Stork Ephippiorhynchus asiaticus prey items in an agricultural landscape|url=https://static1.squarespace.com/static/5c1a9e03f407b482a158da87/t/5c211dc7352f53dd5f7882a9/1545674184080/Black-necked-Stork.pdf|journal=Forktail|volume=27|pages=98–99}}</ref> The foraging method used by the generalists is to stalk or walk across grassland or shallow water, watching for prey.<ref name="HBW" />

===Breeding===

Storks range from being solitary breeders through loose breeding associations to fully [[Bird colony|colonial]]. The jabiru, ''Ephippiorhynchus'' storks and several species of ''Ciconia'' are entirely solitary when breeding.<ref name=":3" /><ref>{{Cite journal|last=Sundar|first=K.S. Gopi|date=2003|title=Notes on the breeding biology of the Black-necked Stork Ephippiorhynchus asiaticus in Etawah and Mainpuri districts, Uttar Pradesh, India|journal=Forktail|volume=19|pages=15–20}}</ref><ref>{{Cite journal|last=Sundar|first=K.S. Gopi|date=2011|title=Agricultural intensification, rainfall patterns, and large waterbird breeding success in the extensively cultivated landscape of Uttar Pradesh, India|url=https://www.sciencedirect.com/science/article/abs/pii/S000632071100365X|journal=Biological Conservation|volume=144|issue=12|pages=3055–3063|doi=10.1016/j.biocon.2011.09.012|bibcode=2011BCons.144.3055G }}</ref> In contrast the ''Mycteria'' storks, Abdim's stork, openbills and ''Leptoptilos'' storks breed in colonies which can range from a couple of pairs to thousands.<ref name=":4" /><ref name=":1" /><ref name=":2" /> Many of these species breed in colonies with other waterbirds, which can include other species of storks, herons and egrets, pelicans, cormorants and ibises. White storks, Oriental storks and Maguari storks are all loosely colonial, and may breed in nests that are within visual range of others of the same species, but have little to do with one another. They also may nest solitarily, and the reasons why they choose to nest together or apart are not understood.<ref name="HBW" /> Storks use trees in a variety of habitats to breed including forests, cities, farmlands, and large wetlands.<ref name=":6" /><ref name=":7" /><ref name=":1" /><ref>{{Cite journal |last=Urfi |first=Abdul J. |date=2010 |title=Using heronry birds to monitor urbanization impacts: a case study of Painted Stork Mycteria leucocephala nesting in the Delhi Zoo, India |journal=Ambio |volume=39 |issue=2 |pages=190–193|doi=10.1007/s13280-010-0018-3 |pmid=20653281 |pmc=3357691 |bibcode=2010Ambio..39..190U }}</ref> 

== In culture ==
[[File:Plate facing page 52, An Argosy of Fables.jpg|thumb|Storks feature in several of [[Aesop's Fables]], such as [[The Fox and the Stork]]]]
Many ancient mythologies feature stories and legends involving storks. In [[Ancient Egypt]], saddle-billed storks were seen as being amongst the most powerful animals and were used to represent the ''[[Ancient Egyptian conception of the soul|ba]]'', the Ancient Egyptian conception of the soul, during the [[Old Kingdom of Egypt|Old Kingdom]].<ref name="Janak2014">{{Cite book |last=Janák |first=Jiří |title=UCLA Encyclopedia of Egyptology |publisher= |year=2014 |editor-last=Wendrich |editor-first=Willeke |location=Los Angeles |pages=1–2, 4 |language=en |chapter=Saddle-billed Stork (ba-bird) |chapter-url=https://escholarship.org/content/qt0r77f2f8/qt0r77f2f8.pdf}}</ref> [[Bennu]], an [[Ancient Egyptian deities|Egyptian deity]] that was later the inspiration for the [[Phoenix (mythology)|phoenix]], may also have been inspired by a stork, although it was more likely an ibis or heron.<ref>{{Cite journal |last=Buckland |first=A. W. |date=1875 |title=Mythological Birds Ethnologically Considered. |url=https://www.jstor.org/stable/pdf/2841232.pdf |journal=The Journal of the Anthropological Institute of Great Britain and Ireland |volume=4 |page=283 |doi=10.2307/2841232|jstor=2841232 }}</ref><ref>{{Cite book |last1=Chadd |first1=Rachel Warren |title=Birds: myth, lore & legend |last2=Taylor |first2=Marianne |date=2016 |publisher=[[Bloomsbury Natural History]] |isbn=978-1-4729-2288-5 |location=London New York, NY |page=19}}</ref>

[[Greek mythology|Greek]] and [[Roman mythology]] portrays storks as models of parental devotion. The 3rd century Roman writer [[Claudius Aelianus|Aelian]], citing the authority of [[Alexander of Myndus]], noted in his ''De natura animalium'' (book 3, chapter 23) that aged storks flew away to oceanic islands where they were transformed into humans as a reward for their piety towards their parents.<ref>{{cite book |url=https://archive.org/stream/L446AelianCharacteristicsOfAnimalsI15/L446-Aelian%20Characteristics%20of%20Animals%20I%3A1-5#page/n104/mode/1up/ |title=Aelian on the characteristics of animals. Volume I. Books I-V. |publisher=Harvard University Press |year=1958 |editor=Schofield, A.F. |page=185}}</ref> Storks were also thought to care for their aged parents, feeding them and even transporting them, and children's books depicted them as a model of filial values. A Greek law called ''Pelargonia'', from the [[Ancient Greek]] word ''pelargos'' for stork, required citizens to take care of their aged parents.{{Cn|date=August 2023}} The Greeks also held that killing a stork could be punished with death.<ref>{{cite book |last=Dolata |first=Pawel T. |title=The White Stork in Poland: Studies in Biology, Ecology and Conservation |publisher=Bogucki Wydawnictwo Naukowe |year=2006 |isbn=978-83-60247-35-8 |editor-last1=Tryjanowski |editor1-first=Piotr |location=Poznań, Poland |chapter=The White Stork ''Ciconia ciconia'' Protection in Poland by Fradition, Customs, Law, and Active Efforts |editor-last2=Sparks |editor-first2=Tim H. |editor-last3=Jerzak |editor-first3=L.}}</ref>

Storks feature in several of Aesop's Fables, most notably in [[The Farmer and the Stork]], [[The Fox and the Stork]], and [[The Frogs Who Desired a King]]. The first fable involves a stork who is caught with a group of cranes who are eating grain in a farmer's field, with the moral that those who associate with wicked people can be held accountable for their crimes.<ref>{{Cite web |title=THE STORK AND THE CRANES |url=http://mythfolklore.net/aesopica/perry/194.htm |access-date=2023-07-30 |website=mythfolklore.net}}</ref> The Fox and the Stork involves a fox who invites a stork for dinner and provides soup in a dish that the stork cannot drink from, and is in turn invited for dinner by the stork and given food in a narrow jug which he cannot access. It cautions readers to follow the principle of [[Primum non nocere|do no harm]].<ref>{{Cite web |title=THE FOX AND THE STORK |url=http://mythfolklore.net/aesopica/perry/426.htm |access-date=2023-07-30 |website=mythfolklore.net}}</ref> The third fable involves a group of frogs that are dissatisfied with the king that Zeus has given them, an inanimate log, and who are then punished with a new [[wiktionary:King_Stork|King Stork]] (a [[Water snake (disambiguation)|water-snake]] in some versions) who eats the frogs.<ref>{{Cite web |title=JUPITER AND THE FROGS |url=http://mythfolklore.net/aesopica/perry/44.htm |access-date=2023-07-30 |website=mythfolklore.net}}</ref><ref>{{Cite book |last=Brewer |first=Ebenezer Cobham |title=Brewer's Dictionary of Phrase & Fable |publisher=Chambers Harrap |year=2012 |isbn=978-0-550-10245-4 |editor-last=Dent |editor-first=Susie |edition=19 |location=London |page=482 |language=en |author-link=Ebenezer Cobham Brewer}}</ref> ''King Stork'' has subsequently entered the English language as a term for a particularly tyrannical ruler.<ref name="OEDKingLog">{{Cite OED|term=King Log|id=4893537697|access-date=2023-07-30}}</ref>

=== Associations with fertility ===
According to European folklore, the white stork is responsible for bringing babies to new parents. The legend is very ancient, but was popularised by an 1839 [[Hans Christian Andersen]] story called "The Storks".<ref name="sax">{{cite book |last=Sax |first=Boria |title=The Mythical Zoo |publisher=ABC-CLIO |year=2001 |isbn=978-1-57607-612-5 |location=Oxford, UK |pages=153–154}}</ref> German folklore held that storks found babies in caves or marshes and brought them to households in a basket on their backs or held in their beaks. These caves contained ''adebarsteine'' or "stork stones". The babies would then be given to the mother or dropped down the chimney. Households would notify when they wanted children by placing sweets for the stork on the window sill.<ref name="Margolis72">{{cite journal |last1=Margolis |first1=Marvin |last2=Parker |first2=Philip |year=1972 |title=The stork fable−some psychodynamic considerations |journal=Journal of the American Psychoanalytic Association |volume=20 |issue=3 |pages=494–511 |doi=10.1177/000306517202000304 |pmid=4116100 |s2cid=45301787}}</ref> Subsequently, the folklore has spread around the world to the [[Philippines]] and countries in [[South America]].<ref name="Margolis72" /> Birthmarks on the back of the head of newborn babies, ''[[nevus flammeus nuchae]]'', are sometimes referred to as stork-bite.<ref>{{cite journal |last1=Jacobs |first1=Alvin H. |last2=Walton |first2=Robert G. |year=1976 |title=The incidence of birthmarks in the neonate |journal=Pediatrics |volume=58 |issue=2 |pages=218–22 |doi=10.1542/peds.58.2.218 |pmid=951136 |s2cid=245028603}}</ref> In [[Slavic mythology|Slavic mythology and pagan religion]], storks were thought to carry unborn [[Soul (spirit)|souls]] from [[Vyraj]] to [[Earth]] in spring and summer.<ref>{{cite book |last=Gieysztor |first=Aleksander |title=Mitologia Słowian |publisher=Wydawnictwa Artystyczne i Filmowe |year=1982 |isbn=978-83-221-0152-0 |location=Warsaw |language=pl |author-link=Aleksander Gieysztor}}</ref> This belief still persists in the modern folk culture of many Slavic countries, in the simplified child story that "storks bring children into the world".<ref>{{cite web |last=Jakubiec |first=Z. |year=2009 |title=Dlaczego bocian przynosił dzieci? |url=http://bocianopedia.pl/bociany-i-bocki/bocian-i-czlowiek-czyli-nasz-powszechnie-lubiany-sasiad/dlaczego-bocian-przynosil-dzieci/157 |access-date=10 March 2011 |work=Bocianopedia |language=pl}}</ref><ref name="autonazwa12">{{cite journal |last=Szczepanowicz |first=Barbara |year=2005 |title=Ptaki Ziemi Świętej: Bocian, czapla, ibis |trans-title=Birds in the Holy Land: Stork, heron, ibis |url=http://basia.tomp.pl/index.php?option=com_content&task=view&id=37&Itemid=12 |journal=Ziemia Święta |language=pl |issue=rok XI 1(41) |access-date=8 February 2011}}</ref>

Famous is the role that the fable played in historical development of [[psychoanalysis]]: the name ‘chimney sweeping’, which the first of all patients gave to her talking cure, is a free association with the place through which the bird used to bring babies into house. Psychoanalyst Marvin Margolis suggests the enduring nature of the stork fable of the newborn is linked to its addressing a psychological need, in that it allays the discomfort of discussing sex and procreation with children. Birds have long been associated with the maternal symbols from pagan goddesses such as [[Juno (mythology)|Juno]], to the [[Holy Spirit in Christianity|Holy Ghost]], and the stork may have been chosen for its white plumage (depicting purity), size, and flight at high altitude (likened to flying between Earth and Heaven).<ref name="Margolis72" />

There were negative aspects to stork folklore as well; a Polish folktale relates how God made the stork's plumage white, while the Devil gave it black wings, imbuing it with both good and evil impulses. They were also associated with handicapped or stillborn babies in Germany, explained as the stork having dropped the baby en route to the household, or as revenge or punishment for past wrongdoing. A mother who was confined to bed around the time of childbirth was said to have been "bitten" by the stork. In Denmark, storks were said to toss a nestling off the nest and then an egg in successive years.<ref name="Margolis72" /> In medieval England, storks were also associated with adultery, possibly inspired by their courtship rituals. Their preening and posture saw them linked with the attribute of self-conceit.<ref name="deVries76">{{cite book |last=de Vries |first=Ad |url=https://archive.org/details/dictionaryofsymb0000vrie/page/445 |title=Dictionary of Symbols and Imagery |publisher=North-Holland Publishing Company |year=1976 |isbn=978-0-7204-8021-4 |location=Amsterdam |page=[https://archive.org/details/dictionaryofsymb0000vrie/page/445 445]}}</ref> Children of [[African American]] [[Slavery in the US|slaves]] were sometimes told that white babies were brought by storks, while black babies were born from [[buzzard]] eggs.<ref>{{cite book |last=Bay |first=Mia |url=https://archive.org/details/isbn_9780195132793 |title=The White Image in the Black Mind: African-American Ideas about White People, 1830–1925 |publisher=[[Oxford University Press]] |year=2000 |isbn=978-0-19-513279-3 |location=New York |page=[https://archive.org/details/isbn_9780195132793/page/120 120] |url-access=registration}}</ref>

=== As food ===
Storks have never been a particularly common food, but occasionally featured in medieval [[Banquet|banquets]].<ref>{{Cite book |last=Davidson |first=Alan |title=The Oxford companion to food |publisher=[[Oxford University Press]] |year=2013 |isbn=978-0-19-280681-9 |editor-last=Jaine |editor-first=Tom |edition=2 |location=Oxford |page=780}}</ref> They may also have been eaten in Ancient Egypt.<ref>{{Cite book |last=Ikram |first=Salima |url=https://books.google.com/books?id=1Am88Yc8gRkC&pg=PR9 |title=Choice cuts: meat production in ancient Egypt |publisher=Peeters |year=1995 |isbn=978-90-6831-745-9 |series=Orientalia Lovaniensia analecta |location=Leuven |pages=25–26}}</ref>

==Footnotes==
{{Reflist|group=note}}

==References==
{{Reflist|30em}}

==External links==
{{wiktionary|Ciconiidae|stork}}
{{Commons category|Ciconiidae}}
{{Wikispecies|Ciconiidae}}
* {{Cite EB1911|wstitle=Stork |short=x}}
* {{Cite Collier's|wstitle=Stork |short=x}}
* [https://web.archive.org/web/20050211205756/http://f01.middlebury.edu/FS010A/STUDENTS/n111.htm Scott MacDonald, "The Stork"] emblematic uses
* [http://www.ebepe.com/html/stork.html Storks] Image documentation
* [http://www.hbw.com/ibc/phtml/familia.phtml?idFamilia=22 Stork videos] on the Internet Bird Collection
* [https://web.archive.org/web/20150126203237/http://jantotrappe.de/stoerche.html Whitestork Photos] Image documentation
* [https://www.youtube.com/watch?v=xhcIvtK-RUA Storks vocalizing]

{{Storks|state=expanded}}
{{Birds}}
{{Taxonbar|from=Q28507}}
{{Authority control}}

[[Category:Ciconiidae| ]]
[[Category:Storks| ]]
[[Category:Extant Rupelian first appearances]]
[[Category:Taxa named by John Edward Gray]]

[[fr:Cigogne]]