Editing Smallmouth bass

{{short description|Species of freshwater fish}}
{{speciesbox
| image = Smallmouth_bass.png
| status = LC
| status_system = IUCN3.1
| status_ref = <ref name="iucn status 19 November 2021">{{cite iucn |author=NatureServe |date=2013 |title=''Micropterus dolomieu'' |volume=2013 |page=e.T202564A18230216 |doi=10.2305/IUCN.UK.2013-1.RLTS.T202564A18230216.en |access-date=19 November 2021}}</ref>
| status2 = G5
| status2_system = TNC
| status2_ref = <ref name="NatureServe">{{cite web|title=Micropterus dolomieu|url=https://explorer.natureserve.org/Taxon/ELEMENT_GLOBAL.2.104786/Micropterus_dolomieu|website=[[NatureServe]] Explorer|access-date=17 April 2024}}</ref>
| taxon = Micropterus dolomieu
| authority = [[Bernard Germain de Lacépède|Lacépède]], 1802
| synonyms = *''Bodianus achigan'' <small>[[Constantine Samuel Rafinesque|Rafinesque]], 1817</small> 
*''Cichla fasciata'' <small>[[Charles Alexandre Lesueur|Lesueur]], 1822</small> 
*''Centrarchus fasciatus'' <small>(Lesueur, 1822)</small> 
*''Cichla minima'' <small>Lesueur, 1822</small>
| synonyms_ref = <ref name = Fishbase>{{Fishbase|Micropterus|dolomieu|month=December|year=2019}}</ref>
}}
The '''smallmouth bass''' ('''''Micropterus dolomieu''''') is a [[species]] of [[freshwater]] [[fish]] in the [[Centrarchidae|sunfish]] [[family (biology)|family]] ([[Centrarchidae]]) of the [[order (biology)|order]] [[Centrarchiformes]].<ref name="fishbase">{{cite web | url=http://www.fishbase.org/Summary/SpeciesSummary.cfm?genusname=Micropterus&speciesname=dolomieu | title=Micropterus dolomieu, Smallmouth bass : Fisheries, gamefish, aquarium }}</ref> It is the [[type species]] of its genus ''[[Micropterus]]'' ([[black bass]]es), and is a popular [[game fish]] sought by anglers throughout the [[temperate zone]]s of [[North America]], and has been spread by [[fish stocking|stocking]] —as well as illegal [[introduced species|introductions]]—to many cool-water tributaries and lakes in Canada and more so introduced in the United States.<ref name="fishbase"/> The maximum recorded size is approximately {{convert|27|in|cm}} and {{convert|12|lb|kg}}. 

The smallmouth bass is native to the upper and middle [[Mississippi River]] basin, the [[Saint Lawrence River]]&ndash;[[Great Lakes]] system, the [[Champlain Valley]], and the [[Hudson Bay]] basin.<ref name="fishbase"/> Its common names include '''smallmouth''', '''bronzeback''', '''brown bass''', '''brownie''', '''smallie''', '''bronze bass''', and '''bareback bass'''.

==Description==
[[File:Smallmouth bass range map.jpg|thumb|Smallmouth Bass Range Map]]
[[File:Detailed underwater photo of smallmouth bass fish micropterus dolomieu.jpg|thumb|right|230px|Detailed underwater photo of Smallmouth Bass ''Micropterus dolomieu''.]]
Smallmouth have a slender but muscular fusiform body shape making them powerful swimmers.<ref name="Wiegmann">Wiegmann, D. D., Baylis, J. R., & Hoff, M. H. (1997, January 01). MALE FITNESS, BODY SIZE AND TIMING OF REPRODUCTION IN SMALLMOUTH BASS, MICROPTERUS DOLOMIEUI. Retrieved from https://esajournals.onlinelibrary.wiley.com/doi/full/10.1890/0012-9658(1997)078[0111:MFBSAT]2.0.CO;2?saml_referrer</ref> The coloration of the smallmouth bass' [[Fish_scale#Ctenoid_scales|ctenoid scales]] range from golden-olive to dark brown [[Dorsal (anatomy)|dorsally]] which fades to a yellowish white [[:File:Anatomical_Directions_and_Axes.JPG|ventrally]] with dark brown vertical bars or blotches along the body and dark brown horizontal bars on the head.<ref name="Wiegmann"/> The combination of the muscular fusiform body shape and camouflage like coloring make these fish highly effective ambush predators. The coloration can vary greatly depending on the fishes age, habitat, water quality, diet, and the spawning cycle.

Generally, the protruding jaw of the smallmouth does not extend back beyond the eyes which are red or brown. They have two dorsal fins which are separated by a shallow interdorsal notch. The front dorsal has 9–11 spiney rays and the back dorsal has 13–15 soft rays.<ref name="fishbase"/>

Males are generally smaller than females. The males tend to range around two pounds, while females can range from three to six pounds. Their average sizes and coloration can differ, depending on if they are found in lacustrine or riverine habitats. Smallmouth found in riverine habitats are generally long and slender which allows greater agility in moving water, while those found in lacustrine habitats and shorter and deeper bodied.<ref name="Schlosser">Schlosser, Isaac J. "The Role of Predation in Age‐ and Size‐Related Habitat Use by Stream Fishes." The Ecological Society of America, John Wiley &amp; Sons, Ltd, 1 June 1987, esajournals.onlinelibrary.wiley.com/doi/abs/10.2307/1938470. </ref>
Riverine smallmouth that live in dark water tend to be rather torpedo-shaped and very dark brown to be more efficient for feeding.<ref name="Schlosser"/> Lacustrine smallmouth bass, however, that live in sandy areas, tend to be a light yellow-brown and are more oval-shaped.<ref name="Schlosser"/>

There are two recognized subspecies, the Northern smallmouth bass (''M. dolomieui dolomieui'') and the Neosho smallmouth bass (''M. dolomieui velox'').<ref name="Missouri-Columbia">University of Missouri-Columbia. (2020, October 19). Management of a popular game fish, the smallmouth bass. ScienceDaily. Retrieved April 26, 2021 from www.sciencedaily.com/releases/2020/10/201019155920.htm
</ref> The Northern smallmouth bass is much more widespread than the much smaller subgroup called the Neosho smallmouth bass. The Neosho are native to an ecologically isolated region of the lower Midwest known as the Central Interior Highlands, which weave through southwestern Missouri, northern Arkansas and northeastern Oklahoma.<ref name="Missouri-Columbia" />

They eat [[tadpole]]s, fish, aquatic insects, and [[crayfish]].<ref>Hanke, Gavin. "Smallmouth Bass." Aliens Among Us. Web. 31 Oct. 2014.</ref><ref>Animal Diversity Web 2008. Micropterus dolomieu: Information. [http://animaldiversity.ummz.mich.edu/site/accounts/information/Micropterus_dolomieu.html]. Accessed 23 April 2021</ref>

The world record size is {{convert|5.41|kg|lboz|order=flip}} caught in the Dale Hollow reservoir, on the [[Kentucky]]—[[Tennessee]] border.<ref>{{Cite web |last=Vitek |first=Jack |date=2015-06-18 |title=10 Biggest Smallmouth Bass World Records of All Time - Game & Fish |url=https://www.gameandfishmag.com/editorial/10-largest-smallmouth-bass-world-records/190547 |access-date=2024-04-19 |website=Game & Fish |language=en}}</ref>

==Habitat==
[[File:Micropterus dolomieu.jpg|thumb|left|Lighter colored variety holding on typical sandy bottom habitat.]]
The smallmouth bass is found in clearer water than the largemouth, especially streams, rivers, and the rocky areas and stumps and also sandy bottoms of lakes and reservoirs. It can also survive in a stronger current than other black bass. The smallmouth prefers cooler water temperatures than its cousin the [[largemouth bass]], and as a result will often seek out deeper, faster moving water during the hot summer months. Because it is intolerant of [[pollution]], the smallmouth bass is a good natural indicator of a healthy environment, though they are still much more resilient than most trout species. Carnivorous, its diet comprises [[crayfish]], [[amphibians]],<ref name="Animal Diversity">{{Cite web|url=https://animaldiversity.org/accounts/Micropterus_dolomieu/|title = Micropterus dolomieu (Bass)| website=[[Animal Diversity Web]] }}</ref> [[insect]]s, and smaller fish, while the larvae feed on various [[zooplankton]]<ref>{{cite web |url=http://academics.cehd.umn.edu/hatch/research/fish/fishes/smallmouth_bass.html |title=Smallmouth Bass |last1=Paulson |first1=Niclole |last2=Hatch |first2=Jay |date=October 23, 2002 |website=University of Minnesota |access-date=June 26, 2019}}</ref> and insect larvae.<ref name="Animal Diversity"/> Adults also cannibalize young of other parents.<ref name="Animal Diversity"/>

The female can lay up to 21,100 eggs, which are guarded by the male in his nest.{{Citation needed|date=May 2012}}

==Migration==
When the weather gets colder, and the water temperature drops below 15 C (60 F), smallmouth will often migrate in search of deeper pools in which they enter a semi-hibernation state,<ref name="dnr.state.mn.us">"Bass biology and identification" see section entitled, "Smallmouth Bass", Minnesota Department of Natural Resources, {{cite web |url=http://www.dnr.state.mn.us/fish/bass/biology.html |title=Bass biology and identification: Minnesota DNR |access-date=2014-07-29 |url-status=dead |archive-url=https://web.archive.org/web/20140811125741/http://www.dnr.state.mn.us/fish/bass/biology.html |archive-date=2014-08-11 }}</ref> moving sluggishly and feeding very little until the warm season returns.<ref name="dnr.state.mn.us"/> The migration patterns of smallmouth have been tracked and it is not unusual for a smallmouth to travel 12 miles in a single day<ref name="dnr.state.mn.us"/> in a stream, creek or river.<ref name="dnr.state.mn.us"/> The overall migration can exceed 60 miles.<ref name="dnr.state.mn.us"/>

Smallmouth generally begin spawning patterns in spring or early summer when water temperatures are between {{convert|15|-|18|C|F|1}}, which is heavily dependent on latitudinal location.<ref name="Ridgway">Ridgway, M., Goff, G., & Miles H. A. Keenleyside. (1989). Courtship and Spawning Behavior in Smallmouth Bass (Micropterus dolomieui). The American Midland Naturalist, 122(2), 209–213. doi:10.2307/2425905</ref> Smallmouth require clean stone, rock, or gravel substrate for a successful spawn.<ref name="Ridgway"/>

==Angling==
{{fly fishing targets}}
{{Main|Bass fishing}}
[[Image:Micropterus dolomieu2.jpg|thumb|200px|Illustration of a group of smallmouth bass]]
[[Image:RainyRiverSmallmouthBass.jpg|thumb|200px|Smallmouth bass from the [[Rainy River (Minnesota–Ontario)|Rainy River]] near [[International Falls, Minnesota]] (Released) ]]
[[Image:SmallMouthBassEagleL.jpg|thumb|200px|Smallmouth bass from [[Eagle Lake (Ontario)|Eagle Lake]] in [[Ontario]], Canada (Released) ]]
In the United States, smallmouth bass were first introduced outside of their native range with the construction of the [[Erie Canal]] in 1825, extending the fish's range into central New York state. During the mid-to-late 19th century, smallmouth were transplanted via the nation's rail system to lakes and rivers throughout the northern and western United States, as far as California. Shippers found that smallmouth bass were a hardy species that could be transported in buckets or barrels by rail, sometimes using the spigots from the railroad water tanks to aerate the fingerlings. They were introduced east of the Appalachians just before the Civil War, and afterwards transplanted to the states of New England.<ref name="Ryan, Will 1996">Ryan, Will, ''Smallmouth Strategies for the Fly Rod'', Lyons & Burford Publishers (1996)</ref><ref name="Waterman">Waterman, Charles F., ''Black Bass & the Fly Rod'', Stackpole Books (1993)</ref>

With increased industrialization and land use changes, many of the nation's eastern trout rivers were polluted or experienced elevated water temperatures, reducing the range of native [[brook trout]]. Smallmouth bass were often introduced to northern rivers with increased water temperatures and slowly became a popular gamefish with many anglers. Equally adaptable to large, cool-water impoundments and reservoirs, the smallmouth also spread far beyond its original native range. Later, smallmouth populations also began to decline after years of damage caused by overdevelopment and pollution, as well as a loss of river habitat caused by damming many formerly wild rivers to form lakes or reservoirs. In recent years, a renewed emphasis on preserving water quality and riparian habitat in the nation's rivers and lakes, together with stricter management practices, eventually benefited smallmouth populations and has caused a resurgence in their popularity with anglers.<ref name="Ryan, Will 1996"/><ref name="Waterman" />

Today, smallmouth bass are very popular [[game fish]], frequently sought by anglers using conventional [[fishing reel|spinning]] and bait casting gear, as well as [[fly fishing]] tackle.<ref name="ReferenceA">Murray, Harry, ''Fly Fishing for Smallmouth Bass'', Lyons Press, 1989</ref><ref name="ReferenceB">Kreh, Lefty, ''Fly Fishing for Bass'', Lyons Press, 2004</ref> The smallmouth bass is potentially the toughest fighting freshwater fish in North America, and is commonly the targeted species in many fresh water fishing tournaments.<ref>Retrieved 2013 - [http://www.fishingtipsdepot.com/bass-fishing-tips.php Bass Fishing Information] - Smallmouth and Largemouth Specific</ref> In addition to wild populations, the smallmouth bass is stocked in cool rivers and lakes throughout Canada and the United States. In shallow streams, it is a wary fish, though usually not to the extent of most trout. The smallmouth is highly regarded for its topwater fighting ability when hooked – old fishing journals referred to the smallmouth bass as "ounce for ounce and pound for pound the gamest fish that swims".<ref>Henshall, James (Dr.), ''Book of the Black Bass'' (1881)</ref> Smallmouth bass are not usually taken for the table, but rather are caught and released by most anglers. However, smaller specimens in cooler water often have higher quality filets of white, firm flesh when cooked.<ref>[http://www.gf.state.az.us/h_f/fish_smallmouth_bass.shtml Arizona Game & Fish Department, ''Smallmouth Bass''] {{webarchive|url=https://web.archive.org/web/20110718130656/http://www.gf.state.az.us/h_f/fish_smallmouth_bass.shtml |date=2011-07-18 }}</ref>

The current all-tackle world record for a smallmouth bass is 11&nbsp;lb 15 oz, caught by David Hayes<ref>{{cite web|url=https://igfa.org/member-services/world-record/common-name/Bass,%20smallmouth|title= IGFA Smallmouth Bass All-Tackle World Record - bass, smallmouth (''Micropterus dolomieu'')|publisher=International Game Fish Association}}</ref> in the [[Dale Hollow Reservoir]], on the Kentucky/Tennessee border, in 1955.<ref>{{cite news |url=https://www.bassmaster.com/news/case-david-hayes|title=60 years ago today: World-record smallmouth caught|website=Bassmaster|date=9 July 2015}}</ref>

===Tackle===
In conventional fishing, smallmouth may be successfully caught on a wide range of natural and artificial baits or lures, including [[Plugs (fishing)|crankbaits]], hair jigs, plastic jerkbaits, artificial worms, [[spinnerbait]]s, and all types of soft plastic lures, including curly tail grubs or tubes with lead head jigs. Spinning reels or baitcasting reels may be used, with line strengths of 6 to 15 pounds typically utilized. According to many, smallmouth typically put up a better, more exciting fight than any other [[black bass]]. Rods are usually of ultralight to medium-heavy action. They may also be caught with a [[fishing rod|fly rod]] using a dry or wet [[artificial fly]], nymphs, streamers, or imitations of larger aquatic creatures, such as [[dobsonfly|hellgrammites]], crawfish, or leeches. Floating topwater popper fly patterns and buzz baits are also popular for smallmouth fishing.<ref name="ReferenceA"/><ref name="ReferenceB"/>

For river fishing, spinning tackle or fly tackle has been the most popular [[angling]] tools for smallmouth in North America for many years. When fishing in colder water, it is believed to be more effective to fish with smaller lures like hair jigs or small spinners. During the rest of the year, smallmouth are usually taken using soft plastic tubes or spinnerbaits. The best spots in rivers to fish for smallmouth are behind rocks or in eddies, where water swirls around. Smallmouth can also be taken in cool lakes like Lake Erie or any of the northern lakes.

==Footnotes==
{{Reflist}}

==References==
{{Wikispecies|Micropterus dolomieu}}
{{Commons}}
* [[ITIS]]: [https://web.archive.org/web/20041031023035/http://www.itis.usda.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=550562 ''Micropterus dolomieu'']
* Rohde, F.&nbsp;C., et al. ''Freshwater Fishes of the Carolinas, Virginia, Maryland, and Delaware.'' Chapel Hill: University of North Carolina Press, 1994.
* Whitlock, John, "[http://animaldiversity.ummz.umich.edu/site/accounts/information/Micropterus_dolomieu.html ''Micropterus dolomieu: Information'']". ''Animal Diversity Web.'' Ann Arbor: University of Michigan Museum of Zoology, 2004.
* New River in Virginia
* {{cite book |last=Jones |first=Sheridan R. |title=Black Bass & Bass-Craft: The Life and Habits of the Two Bass and Successful Angling Strategy |year=1924 |publisher=MacMillan |location=New York }}
*{{cite book |last=Ovington |first=Ray |title=Tactics on Bass--How to Wade, Cast, and Fish Out Each of 23 Different Kinds of Bass Areas |year=1983 |publisher=Charles Scribner's & Sons |location=New York |isbn=0-684-17860-5 |url-access=registration |url=https://archive.org/details/tacticsonbass00ovin }}
*Gunn, J.C., Berkman, L.K., Koppelman, J. et al. Complex patterns of genetic and morphological differentiation in the Smallmouth Bass subspecies (Micropterus dolomieu dolomieu and M. d. velox) of the Central Interior Highlands. Conserv Genet 21, 891–904 (2020). https://doi.org/10.1007/s10592-020-01295-1
*Effects of temperature and Streamflow on time and duration of spawning by Smallmouth bass. (2011, January 9). Taylor & Francis. https://www.tandfonline.com/doi/abs/10.1577/1548-8659(1986)115%3C693%3AEOTASO%3E2.0.CO%3B2 
*U.S. Fish and Wildlife Service/Fish and Aquatic Conservation. (n.d.). Smallmouth bass. Official Web page of the U S Fish and Wildlife Service. https://www.fws.gov/fisheries/freshwater-fish-of-america/smallmouth_bass.html
{{Taxonbar|from=Q536224}}

[[Category:Micropterus|Smallmouth bass]]
[[Category:Freshwater fish of the United States]]
[[Category:Fish of the Eastern United States]]
[[Category:Fish of the Great Lakes]]
[[Category:Fauna of the Northeastern United States]]
[[Category:Fauna of the Southeastern United States]]
[[Category:Freshwater fish of the Southeastern United States]]
[[Category:Fish described in 1802|Smallmouth bass]]
[[Category:Taxa named by Bernard Germain de Lacépède]]