Editing Quercus velutina

{{Short description|Species of oak tree}}
{{Speciesbox
|name=Black oak
|image=Quercus velutina 001.jpg
|image_caption=Planted tree
|status=LC
|status_system=IUCN3.1
|status_ref=<ref>{{cite iucn |author=Wenzell, K. |author2=Kenny, L. |year=2015 |title=Quercus velutina |volume=2015 |page=e.T194244A2305832 |doi=10.2305/IUCN.UK.2015-4.RLTS.T194244A2305832.en |access-date=12 April 2020}}</ref>
|genus=Quercus
|display_parents=2
|parent=Quercus sect. Lobatae
|species=velutina
|authority=[[Jean-Baptiste Lamarck|Lam.]]<ref>{{cite encyclopedia |last=Lamarck |first=Jean-Baptiste Pierre Antoine de Monet de |date=1785 |title=[[Encyclopédie méthodique]]. Botanique |volume=1 |page=[https://biodiversitylibrary.org/page/717313 721]}}</ref>
|range_map=Quercus velutina range map 1.png
|range_map_caption=Generalized natural range
|synonyms_ref=<ref>{{ThePlantList |authority=Lam. |access-date=2016-04-27}}</ref>
|synonyms={{collapsible list|bullets=true
  |''Quercus discolor'' <small>Aiton</small>
  |''Quercus leiodermis'' <small>Ashe</small>
  |''Quercus missouriensis'' <small>Ashe</small>
  |''Quercus tinctoria'' <small>Bartram</small>
  |''Quercus tinctoria'' <small>Michx.</small>
  }}
}}

'''''Quercus velutina''''' (Latin 'velutina', "velvety") , the '''black oak''', is a species of [[oak]] in the [[red oak]] group (''Quercus'' sect. ''Lobatae''), native and widespread in eastern and central North America. It is sometimes called the '''eastern black oak'''.{{r|brandeis}}

''Quercus velutina'' was previously known as yellow oak due to the yellow pigment in its inner bark. It is a close relative of the [[California black oak]] (''Quercus kelloggii'') found in western North America.

==Description==
In the northern part of its range, ''Quercus velutina'' is a relatively small tree, reaching a height of {{Convert|20-25|m|abbr=off}} and a diameter of {{Convert|90|cm|abbr=off}}, but it grows larger in the south and center of its range, where heights of up to {{cvt|42|m|ft}} are known.

The [[leaves]] of the black oak are alternately arranged on the twig and are {{cvt|10|–|20|cm|in|0}} long with 5–7 bristle-tipped lobes separated by deep U-shaped notches. The upper surface of the leaf is a shiny deep green, and the lower is yellowish-brown. There are also stellate hairs on the underside of the leaf that grow in clumps.<ref name=FNA /> Some key characteristics for identification include that leaves grown in the sun have very deep U-shaped sinuses and that the buds are velvety and covered in white hairs.{{r|FNA|vt}}

Black oak is [[monoecious]]. The [[staminate]] flowers develop from leaf axils of the previous year and the [[catkin]]s emerge before or at the same time as the current leaves in April or May. The [[pistillate]] flowers are borne in the [[axil]]s of the current year's leaves and may be solitary or occur in two- to many-flowered spikes. The fruit, an [[acorn]] that occurs singly or in clusters of two to five, is about one-third enclosed in a scaly cup and matures in 2 years. Black oak acorns are brown when mature and ripen from late August to late October, depending on geographic location.

The [[fruit]]s or [[acorn]]s of the black oak are medium-sized and broadly rounded.<ref>{{cite book |last1=Zim |first1=Herbert S. |last2=Martin |first2=Alexander C. |date=1987 |title=Trees: A Guide to Familiar American Trees |series=[[Golden Guide]]s |publisher=Golden Books |isbn=9780307240569 |url-access=registration |url=https://archive.org/details/trees00herb }}</ref> The cap is large and covers almost half of the nut.<ref name=FNA />

{{Gallery
|Bark black oak 8771.jpg|Detail of mature bark
|Quercus velutina leaves.jpg|Young leaves are densely pubescent (hairy)
|Quercus velutina (23537664393).jpg|Winter twig
|Quercus velutina (23837102639).jpg|Acorns
}}

==Habitat and distribution==
{{Unreferenced section|date=August 2020}}
Black oak is found in all the coastal states from [[Maine]] to [[Texas]], inland as far as [[Michigan]], [[Ontario]], [[Minnesota]], [[Nebraska]], [[Kansas]], [[Oklahoma]], and eastern [[Texas]].<ref name="bonap">{{BONAP|ref|genus=Quercus|species=velutina|access-date=27 April 2016}}</ref>

It grows on all aspects and slope positions. It grows best in coves and on middle and lower slopes with northerly and easterly aspects. It is found at elevations up to {{cvt|1200|m|ft}} in the southern [[Appalachians]]. In southern New England, black oak grows on cool, moist soils. Elsewhere it occurs on warm, moist soils.
The most widespread soils on which black oak grows are the [[Alfisol|udalfs]] and [[Mollisol|udolls]]. These soils are derived from glacial materials, sandstones, shales, and limestone and range from heavy clays to loamy sands with some having a high content of rock or chert fragments. Black oak grows best on well drained, silty clay to loam soils.

The most important factors determining site quality for black oak are the thickness and texture of the [[Soil horizon|A horizon]], texture of the [[Soil horizon|B horizon]], [[Aspect (geography)|aspect]], and slope position. Other factors may be important in localized areas. For example, in northwestern West Virginia increasing precipitation to {{convert|1120|mm|in}} resulted in increased site quality; more than {{cvt|1120|mm|in}} had no further effect. In southern Indiana, decreasing site quality was associated with increasing slope steepness.

Near the limits of its range, [[Topography|topographic]] factors may restrict its distribution. At the western limits black oak is often found only on north and east aspects where moisture conditions are most favorable. In southern Minnesota and Wisconsin it is usually found only on ridge tops and the lower two-thirds of south- and west-facing slopes.

==Ecology==

=== Associated plant species ===
Common tree associates of black oak are white oak (''[[Quercus alba]]''), northern red oak (''[[Quercus rubra]]''), pignut hickory (''[[Carya glabra]]''), [[mockernut hickory]] (''C.&nbsp;tomentosa''), [[bitternut hickory]] (''C.&nbsp;cordiformis''), and [[shagbark hickory]] (''C.&nbsp;ovata''); American elm (''[[Ulmus americana]]'') and [[slippery elm]] (''U.&nbsp;rubra''); white ash (''[[Fraxinus americana]]''); black walnut (''[[Juglans nigra]]'') and [[butternut tree|butternut]] (''J.&nbsp;cinerea''); scarlet oak (''[[Quercus coccinea]]''), [[southern red oak]] (''Q.&nbsp;falcata''), and [[chinkapin oak]] (''Q.&nbsp;muehlenbergii''); red maple (''[[Acer rubrum]]'') and [[sugar maple]] (''A.&nbsp;saccharum''); black cherry (''[[Prunus serotina]]''); and blackgum (''[[Nyssa sylvatica]]'').{{citation needed|date=August 2020}}

Common small tree associates of black oak include flowering dogwood (''[[Cornus florida]]''), sourwood (''[[Oxydendrum arboreum]]''), sassafras (''[[Sassafras albidum]]''), eastern hophornbeam (''[[Ostrya virginiana]]''), redbud (''[[Cercis canadensis]]''), pawpaw (''[[Asimina triloba]]''), downy serviceberry (''[[Amelanchier arborea]]''), and American bladdernut (''[[Staphylea trifolia]]''). Common shrubs include ''[[Vaccinium]]'' spp., mountain-laurel (''[[Kalmia latifolia]]''), witch-hazel (''[[Hamamelis virginiana]]''), beaked hazel (''[[Corylus cornuta]]''), spicebush (''[[Lindera benzoin]]''), [[sumac]] (''Rhus'' spp.), and ''[[Viburnum]]'' spp. The most common vines are greenbrier (''[[Smilax]]'' spp.), grape (''[[Vitis]]'' spp.), poison-ivy (''[[Toxicodendron radicans]]''), and Virginia creeper (''[[Parthenocissus quinquefolia]]'').{{citation needed|date=August 2020}}

Black oak is often a predominant species in the canopy of an [[oak–heath forest]].<ref>[https://web.archive.org/web/20090115181617/http://www.dcr.virginia.gov/natural_heritage/ncTIIIe.shtml ''The Natural Communities of Virginia Classification of Ecological Community Groups'']. (Version 2.3), Virginia Department of Conservation and Recreation, 2010</ref><ref>Schafale, M. P. and Weakley, A. S. (1990). ''Classification of the natural communities of North Carolina: third approximation''. North Carolina Natural Heritage Program, North Carolina Division of Parks and Recreation.</ref>

=== Seed production and dissemination ===
{{Unreferenced section|date=August 2020}}
In forest stands, black oak begins to produce seeds at about age 20 and reaches optimum production at 40 to 75 years. It is a consistent seed producer with good crops of acorns every 2 to 3 years. In Missouri, the average number of mature acorns per tree was generally higher than for other oaks over a 5-year period, but the number of acorns differed greatly from year to year and from tree to tree within the same stand.

The number of seeds that become available for regenerating black oak may be low even in good seed years. Insects, squirrels, deer, turkey, small rodents, and birds consume many [[acorn]]s. They can eat or damage a high percentage of the acorn crop in most years and essentially all of it in poor seed years.

Black oak acorns from a single tree are dispersed over a limited area by squirrels, mice, and gravity. The blue jay may disperse over longer distances.

=== Response to competition ===
{{Unreferenced section|date=August 2020}}
Black oak is classed as intermediate in tolerance to shade. It is less tolerant than many of its associates such as white and chestnut oaks, hickories, beech (''[[Fagus grandifolia]]''), maples, elm, and blackgum. However, it is more tolerant than yellow-poplar (''Liriodendron tulipifera''), black cherry, and shortleaf pine (''[[Pinus echinata]]''). It is about the same as northern red oak and scarlet oak. Seedlings usually die within a few years after being established under fully stocked over stories. Most black oak sprouts under mature stands develop crooked stems and flat-topped or misshapen crowns. After the over story is removed, only the large stems are capable of competing successfully. Seedlings are soon overtopped. The few that survive usually remain in the intermediate crown class.

Even-aged silvicultural systems satisfy the reproduction and growth requirements of black oak better than the all-aged or uneven-aged selection system. Under the selection system, black oak is unable to reproduce because of inadequate light. Stands containing black oak that are managed under the selection system will gradually be dominated by more shade-tolerant species.

Dormant buds are numerous on the boles of black oak trees. These buds may be stimulated to sprout and produce branches by mechanical pruning or by exposure to greatly increased light, as by thinning heavily or creating openings in the stand. Dominant trees are less likely to produce epicormic branches than those in the lower crown class.

=== Damaging agents ===
{{Unreferenced section|date=August 2020}}
Wildfires seriously damage black oak trees by killing the [[cambium]] at the base of the trees. This creates an entry point for decay fungi. The result is loss of volume because of heart rot. Trees up to pole size are easily killed by fire and severe fires may even kill saw timber. Many of the killed trees sprout and form a new stand. However, the economic loss may be large unless at least some of it can be salvaged.

Oak wilt (''[[Bretziella fagacearum]]'') is a potentially serious vascular disease of black oak that is widespread throughout the eastern United States. Trees die within a few weeks after the symptoms first appear. Usually scattered individuals or small groups of trees are killed, but areas several hectares in size may be affected. The disease is spread from tree to tree through root grafts and over larger distances by sap-feeding beetles ([[Nitidulidae]]) and the small oak bark beetle.

Shoestring root rot (''[[Armillaria mellea]]'') attacks black oak and may kill trees weakened by fire, lightning, drought, insects, or other diseases. A root rot, ''[[Phytophthora cinnamomi]]'', may kill seedlings in the nursery. Cankers caused by ''[[Strumella (moth)|Strumella]]'' and ''[[Nectria]]'' species damage the holes of black oak but seldom kill trees. Foliage diseases that attack black oak are the same as those that typically attack species in the red oak group and include anthracnose (''[[Gnomonia quercina]]''), leaf blister (''[[Taphrina]]'' spp.), powdery mildews (''[[Phyllactinia corylea]]'' and ''[[Microsphaera alni]]''), oak-pine rusts (''[[Cronartium]]'' spp.), and leaf spots (''[[Actinopelte dryina]]'').

Tunneling insects that attack the boles of black oak and cause serious lumber degrade include the carpenter worm (''[[Prionoxystus robiniae]]''), red oak borer (''[[Enaphalodes rufulus]]''), the twolined chestnut borer (''[[Agrilus bilineatus]]''), the oak timber worm (''[[Arrenodes minutus]]''), and the Columbian timber beetle (''[[Corthylus columbianus]]'').

The gypsy moth (''[[Lymantria dispar]]'') feeds on foliage and is potentially the most destructive insect. Although black oaks withstood a single defoliation, two or three defoliations in successive years kill many trees. Other defoliators that attack black oak and may occasionally be epidemic are the variable oak leaf caterpillar (''[[Heterocampa manteo]]''), the orange striped oakworm (''[[Anisota senatoria]]''), and the brown tail moth (''[[Euproctis chrysorrhoea]]'').

The nut weevils (''[[Curculio]]'' spp.), gall-forming cynipids (''[[Callirhytis]]'' spp.), filbertworm (''[[Melissopus latiferreanus]]''), and acorn moth (''[[Valentinia glandulella]]'') damage black oak acorns.

==Named hybrids involving black oak==
Black oak is well known to readily hybridize with other members of the [[red oak]] (''Quercus'' sect. ''Lobatae'') group, being one parent in at least a dozen different named hybrids.{{r|FNA}}

*[[Quercus × bushii|''Quercus'' × ''bushii'']] (''[[Quercus marilandica]]'' × ''Q.&nbsp;velutina'') – Bush's oak
*[[Quercus × cocksii|''Quercus'' × ''cocksii'']] (''[[Quercus laurifolia]]'' × ''Q.&nbsp;velutina'') – Cocks' oak
*[[Quercus × demarei|''Quercus'' × ''demarei'']] (''[[Quercus nigra]]'' × ''Q.&nbsp;velutina'')
*[[Quercus × discreta|''Quercus'' × ''discreta'']] (''[[Quercus shumardii]]'' × ''Q.&nbsp;velutina'')
*[[Quercus × filialis|''Quercus'' × ''filialis'']] (''[[Quercus phellos]]'' × ''Q.&nbsp;velutina'')
*[[Quercus × fontana|''Quercus'' × ''fontana'']] (''[[Quercus coccinea]]'' × ''Q.&nbsp;velutina'')
*[[Quercus × hawkinsiae|''Quercus'' × ''hawkinsiae'']] (''[[Quercus rubra]]'' × ''Q.&nbsp;velutina'') – Hawkin's oak
*[[Quercus × leana|''Quercus'' × ''leana'']] (''[[Quercus imbricaria]]'' × ''Q.&nbsp;velutina'') – Lea's oak
*[[Quercus × palaeolithicola|''Quercus'' × ''palaeolithicola'']] (''[[Quercus ellipsoidalis]]'' × ''Q.&nbsp;velutina'')
*[[Quercus × podophylla|''Quercus'' × ''podophylla'']] (''[[Quercus incana]]'' × ''Q.&nbsp;velutina'')
*[[Quercus × rehderi|''Quercus'' × ''rehderi'']] (''[[Quercus ilicifolia]]'' × ''Q.&nbsp;velutina'') – Rehder's oak
*[[Quercus × vaga|''Quercus'' × ''vaga'']] (''[[Quercus palustris]]'' × ''Q.&nbsp;velutina'')
*[[Quercus × willdenowiana|''Quercus'' × ''willdenowiana'']] (''[[Quercus falcata]]'' × ''Q.&nbsp;velutina'') – Willdenow's oak

==Uses==
The inner [[Bark (botany)|bark]] of the black oak contains a yellow-orange coloring from the pigment [[quercitron]], which was sold commercially in Europe until the 1940s, and lending the species its former common name of yellow oak.<ref>{{Silvics|first=Ivan L.|last=Sander|volume=2|genus=Quercus|species=velutina|access-date=18 November 2012}}</ref>

==References==
{{Reflist|refs=
<ref name=brandeis>{{cite web |title=Eastern Black Oak |url=http://www.bio.brandeis.edu/fieldbio/Clapp_Hansen_Siegel/Pages/Black_Oak.html |website=www.bio.brandeis.edu}}</ref>
<ref name=FNA>{{eFloras|1|233501095|Quercus velutina |family=Fagaceae |first=Kevin C. |last=Nixon |access-date=27 April 2016}}</ref>
<ref name=vt>[https://web.archive.org/web/20080430085649/http://www.cnr.vt.edu/DENDRO/dendrology/syllabus/factsheet.cfm?ID=39 Virginia Tech Department of Forestry fact sheet on ''Black Oak'']</ref>
}}

{{Taxonbar|from=Q1001011}}
{{Authority control}}

[[Category:Quercus|velutina]]
[[Category:Trees of Northern America]]
[[Category:Hardwood forest plants]]
[[Category:Plant dyes]]
[[Category:Trees of humid continental climate]]
[[Category:Plants described in 1785]]