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{{Short description|Class of tardigrades}} {{Automatic taxobox | image = Echiniscus succineus (10.3897-evolsyst.3.33580) Figure 2.jpg | image_caption = ''Echiniscus succineus'' under [[Phase-contrast microscopy|PCM]] (left) and [[Scanning electron microscope|SEM]] (right) | taxon = Heterotardigrada | authority = [[Ernst Marcus (zoologist)|Marcus]], 1927 | subdivision_ranks = Families | subdivision = '''Order [[Arthrotardigrada]]'''<br/> [[Archechiniscidae]]<br/> [[Batillipedidae]]<br/> [[Coronarctidae]]<br/> [[Halechiniscidae]]<br/> [[Neoarctidae]]<br /> [[Neostygarctidae]]<br /> [[Renaudarctidae]]<br/> [[Stygarctidae]]<br/> [[Styraconyxidae]]<br/> [[Tanarctidae]]<br /> '''Order [[Echiniscoidea]]'''<br/> [[Carphaniidae]]<br/> [[Echiniscidae]]<br/> [[Echiniscoididae]]<br/> [[Oreellidae]] }} '''Heterotardigrades''' (class '''Heterotardigrada''') is a [[class (biology)|class]] of the [[tardigrade]]s (water bears) that have cephalic appendages and legs with four separate but similar digits or claws on each. 444 species have been described.<ref name=Zhang2011>{{cite journal| author=Zhang, Z.-Q.| title=Animal biodiversity: An introduction to higher-level classification and taxonomic richness | journal=Zootaxa| volume=3148| year=2011| pages=7β12| doi=10.11646/zootaxa.3148.1.3 | url=http://mapress.com/zootaxa/2011/f/zt03148p012.pdf}}</ref> == Anatomy == The anatomy of the reproductive system is an important defining feature in distinguishing the different groups of tardigrades. Heterotardigrades have gonoducts that open to the outside through a preanal gonopore, rather than opening into the rectum as in the [[Eutardigrada]]. == Ecology == Some [[order (biology)|order]]s of heterotardigrades are marine, others are terrestrial, but as for all tardigrades, all are aquatic in the sense that they must be surrounded by at least a film of moisture in order to be active β though they can survive in a dormant state if the habitat dries out. == References == {{Reflist}} == External links == {{Commons category|Heterotardigrada}} {{Wikispecies|Heterotardigrada}} * {{ITIS |id=155167 |taxon=Heterotardigrada}} * {{ITIS |id=155168 |taxon=Arthrotardigrada}} * {{ITIS |id=155299 |taxon=Echiniscoidea}} {{Tardigrades}} {{Taxonbar|from=Q138557}} {{Authority control}} [[Category:Heterotardigrada| ]] [[Category:Ecdysozoa classes]] [[Category:Taxa named by Ernst Marcus (zoologist)]]
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