Editing Pupil (section)

=== Activity pattern and behavior ===
In a study of [[Australia]]n [[snake]]s, pupil shapes correlated both with [[diel]] activity times and with foraging behavior. Most snake species with vertical pupils were [[nocturnal]] and also ambush foragers, and most snakes with circular pupils were [[diurnality|diurnal]] and active foragers. Overall, foraging behaviour predicted pupil shape accurately in more cases than did diel time of activity, because many active-foraging snakes with circular pupils were not diurnal.  It has been suggested that there may be a similar link between foraging behaviour and pupil shape amongst the felidae and canidae discussed above.<ref name="Brischoux et al., (2010)" />

A 2015 study<ref>{{cite journal |last1=Banks |first1=Martin S.  |last2=Sprague |first2=William W.  |last3=Schmoll |first3=Jürgen |last4=Parnell |first4=Jared A. Q.  |last5=Love |first5=Gordon D. |title=Why do animal eyes have pupils of different shapes? |journal=Science Advances |volume=1 |issue=7 |pages=e1500391 |year=2015 |doi=10.1126/sciadv.1500391 |pmid=26601232 |pmc=4643806 |bibcode=2015SciA....1E0391B |doi-access=free }}</ref> confirmed the hypothesis that elongated pupils have increased dynamic range, and furthered the correlations with diel activity. However it noted that other hypotheses could not explain the orientation of the pupils. They showed that vertical pupils enable ambush predators to optimise their depth perception, and horizontal pupils to optimise the field of view and image quality of horizontal contours. They further explained why elongated pupils are correlated with the animal's height.

<gallery widths="240" heights="180" caption="Animals with non-circular pupils">
File:Horizontal Rectangular Pupil of a Goat.jpg|A [[goat]] with horizontal rectangular pupils
File:Taeniura_grabata_eye.jpg|A [[stingray]] with crescent pupils
File:Crocodylus siamensis closeup.jpg|A [[crocodile]] with thin vertical slit pupils
File:Cuttlefish_eye.jpg|A [[cuttlefish]] with W-shaped pupils
File:Gecko-oeil.jpg|A [[gecko]] with 'thin string of pearls' pupils
File:Catpupil03042006.jpg|A [[cat]] with thick vertical slit pupils
</gallery>