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===Epidermis=== The [[Epidermis (botany)|plant epidermis]] is specialised tissue, composed of parenchyma cells, that covers the external surfaces of leaves, stems and roots. Several cell types may be present in the epidermis. Notable among these are the stomatal guard cells that control the rate of [[Gas exchange#Plants|gas exchange]] between the plant and the atmosphere, glandular and clothing hairs or [[trichome]]s, and the [[root hair]]s of primary roots. In the shoot epidermis of most plants, only the [[stomata|guard cells]] have chloroplasts. Chloroplasts contain the green pigment chlorophyll which is needed for photosynthesis. The epidermal cells of aerial organs arise from the superficial layer of cells known as the ''tunica'' (L1 and L2 layers) that covers the plant [[meristem|shoot apex]],<ref name="Cutter"/> whereas the cortex and vascular tissues arise from innermost layer of the shoot apex known as the ''corpus'' (L3 layer). The epidermis of roots originates from the layer of cells immediately beneath the root cap. The epidermis of all aerial organs, but not roots, is covered with a [[plant cuticle|cuticle]] made of [[polyester]] [[cutin]] or polymer [[Cutan (polymer)|cutan]] (or both), with a superficial layer of [[epicuticular wax]]es. The epidermal cells of the primary shoot are thought to be the only plant cells with the biochemical capacity to synthesize cutin.<ref name="Kolattukudy 1996">Kolattukudy, PE (1996) Biosynthetic pathways of cutin and waxes, and their sensitivity to [[environmental stresses]]. In: Plant Cuticles. Ed. by G. Kerstiens, BIOS Scientific publishers Ltd., Oxford, pp 83β108</ref>
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