Editing Centriole (section)

==Origin==
[[Eukaryogenesis|LECA]], the last common ancestor of all [[eukaryote]]s was a [[cilia]]ted cell with centrioles.{{Citation needed|date=April 2024}} Some lineages of eukaryotes, such as [[land plants]], do not have centrioles except in their motile male gametes. Centrioles are completely absent from all cells of [[Pinophyta|conifers]] and [[angiosperm|flowering plants]], which do not have ciliate or flagellate gametes.<ref>{{cite journal | last1 = Marshall | first1 = W.F. | year = 2009 | title = Centriole Evolution | journal = Current Opinion in Cell Biology | volume = 21 | issue = 1| pages = 14–19 | doi = 10.1016/j.ceb.2009.01.008 | pmid=19196504 | pmc=2835302}}</ref>
It is unclear if the last common ancestor had one<ref name="Bornens-2007">{{Cite book| doi = 10.1007/978-0-387-74021-8_10| last2 = Azimzadeh| pmid = 17977464| isbn = 978-0-387-74020-1| year = 2007| pages = [https://archive.org/details/eukaryoticmembra00gasp/page/119 119–129]| series = Advances in Experimental Medicine and Biology| last1 = Bornens| first2 = J.| chapter = Origin and Evolution of the Centrosome| title = Eukaryotic Membranes and Cytoskeleton| volume = 607| first1 = M.| chapter-url-access = registration| chapter-url = https://archive.org/details/eukaryoticmembra00gasp| url = https://archive.org/details/eukaryoticmembra00gasp/page/119}}</ref> or two cilia.<ref>{{Cite journal | doi = 10.1093/gbe/evp011 | last1 = Rogozin | first1 = I. B. | last2 = Basu | first2 = M. K. | last3 = Csürös | first3 = M. | last4 = Koonin | first4 = E. V. | title = Analysis of Rare Genomic Changes Does Not Support the Unikont-Bikont Phylogeny and Suggests Cyanobacterial Symbiosis as the Point of Primary Radiation of Eukaryotes | journal = Genome Biology and Evolution | volume = 1 | pages = 99–113 | year = 2009 | pmid = 20333181 | pmc = 2817406}}</ref> Important genes such as those coding for [[centrins]], required for centriole growth, are only found in eukaryotes, and not in [[bacteria]] or [[archaea]].<ref name="Bornens-2007"/>