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===Genetics and environment=== Both genetics and environment have been implicated in influencing pro-social or altruistic behavior.<ref>{{cite book |doi=10.1093/acprof:oso/9780199964772.003.0001 |title=Prosocial Development |date=2014 |publisher=Oxford University Press |isbn=978-0-19-996477-2 |editor-last1=Padilla-Walker |editor-last2=Carlo |editor-first1=Laura M. |editor-first2=Gustavo |chapter=1 The Study of Prosocial Behavior: Past, Present, and Future |pages=3β16 }}</ref> Candidate genes include OXTR ([[gene polymorphism|polymorphism]]s in the [[oxytocin receptor]]),<ref>{{cite journal |last1=Singer |first1=Tania |last2=Snozzi |first2=Romana |last3=Bird |first3=Geoffrey |last4=Petrovic |first4=Predrag |last5=Silani |first5=Giorgia |last6=Heinrichs |first6=Markus |last7=Dolan |first7=Raymond J. |title=Effects of oxytocin and prosocial behavior on brain responses to direct and vicariously experienced pain |journal=Emotion |date=December 2008 |volume=8 |issue=6 |pages=781β791 |doi=10.1037/a0014195 |pmid=19102589 |pmc=2672051 }}</ref><ref>{{cite journal |last1=Israel |first1=Salomon |last2=Weisel |first2=Ori |last3=Ebstein |first3=Richard P. |last4=Bornstein |first4=Gary |title=Oxytocin, but not vasopressin, increases both parochial and universal altruism |journal=Psychoneuroendocrinology |date=August 2012 |volume=37 |issue=8 |pages=1341β1344 |doi=10.1016/j.psyneuen.2012.02.001 |pmid=22377540 }}</ref><ref>{{cite journal |last1=Barraza |first1=Jorge A. |last2=McCullough |first2=Michael E. |last3=Ahmadi |first3=Sheila |last4=Zak |first4=Paul J. |title=Oxytocin infusion increases charitable donations regardless of monetary resources |journal=Hormones and Behavior |date=July 2011 |volume=60 |issue=2 |pages=148β151 |doi=10.1016/j.yhbeh.2011.04.008 |pmid=21596046 }}</ref> [[CD38]], [[COMT]], [[DRD4]], [[DRD5]], [[IGF2]], [[AVPR1A]]<ref>{{cite journal | doi=10.1371/journal.pone.0025274 | doi-access=free | title=AVPR1A Variant Associated with Preschoolers' Lower Altruistic Behavior | date=2011 | last1=Avinun | first1=Reut | last2=Israel | first2=Salomon | last3=Shalev | first3=Idan | last4=Gritsenko | first4=Inga | last5=Bornstein | first5=Gary | last6=Ebstein | first6=Richard P. | last7=Knafo | first7=Ariel | journal=PLOS ONE | volume=6 | issue=9 | pages=e25274 | pmid=21980412 | pmc=3182215 | bibcode=2011PLoSO...625274A }}</ref> and [[GABRB2]].<ref>{{cite journal| pmc=3871336 | pmid=24132092 | doi=10.1098/rsbl.2013.0395 | volume=9 | issue=6 | title=Genes underlying altruism | year=2013 | journal=Biol Lett | page=20130395 | last1 = Thompson | first1 = GJ | last2 = Hurd | first2 = PL | last3 = Crespi | first3 = BJ}}</ref> It is theorized that some of these genes influence altruistic behavior by modulating levels of neurotransmitters such as [[serotonin]] and [[dopamine]]. According to [[Christopher Boehm]], altruistic behaviour evolved as a way of surviving within a group.<ref>{{cite journal | url=https://psycnet.apa.org/record/2011-29744-000 | title=Moral origins: The evolution of virtue, altruism, and shame | journal=Apa Psycnet | date=2012 | last1=Christopher | first1=Boehm }}</ref>
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