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=== Algae === One type of [[organelle]], originating from [[cyanobacteria]]l [[endosymbiont]]s called [[UCYN-A]]2,<ref name="Thompson_2012" /><ref>{{Cite journal |last1=Thompson |first1=Anne |last2=Carter |first2=Brandon J. |last3=Turk-Kubo |first3=Kendra |last4=Malfatti |first4=Francesca |last5=Azam |first5=Farooq |last6=Zehr |first6=Jonathan P. |date=October 2014 |title=Genetic diversity of the unicellular nitrogen-fixing cyanobacteria UCYN-A and its prymnesiophyte host: UCYN-A genetic diversity |url=https://cloudfront.escholarship.org/dist/prd/content/qt4687q7k8/qt4687q7k8.pdf?t=nx0365 |journal=Environmental Microbiology |language=en |volume=16 |issue=10 |pages=3238–3249 |doi=10.1111/1462-2920.12490 |pmid=24761991 |s2cid=24822220}}</ref> can turn nitrogen gas into a biologically available form. This [[nitroplast]] was discovered in [[algae]], particularly in the marine algae [[Braarudosphaera bigelowii]].<ref>{{Cite journal |last=Wong |first=Carissa |date=2024-04-11 |title=Scientists discover first algae that can fix nitrogen — thanks to a tiny cell structure |url=https://www.nature.com/articles/d41586-024-01046-z |journal=Nature |volume=628 |issue=8009 |page=702 |language=en |doi=10.1038/d41586-024-01046-z|pmid=38605201 |bibcode=2024Natur.628..702W }}</ref> [[Diatom]]s in the family ''Rhopalodiaceae'' also possess [[cyanobacteria]]l [[endosymbiont]]s called spheroid bodies or diazoplasts.<ref>{{cite journal |last1=Moulin |first1=Solène L. Y. |last2=Frail |first2=Sarah |last3=Braukmann |first3=Thomas |last4=Doenier |first4=Jon |last5=Steele-Ogus |first5=Melissa |last6=Marks |first6=Jane C. |last7=Mills |first7=Matthew M. |last8=Yeh |first8=Ellen |date=15 April 2024 |title=The endosymbiont of Epithemia clementina is specialized for nitrogen fixation within a photosynthetic eukaryote |journal=ISME Communications |volume=4 |pages=ycae055 |doi=10.1093/ismeco/ycae055 |pmc=11070190 |pmid=38707843 |doi-access=free}}</ref> These endosymbionts have lost photosynthetic properties, but have kept the ability to perform nitrogen fixation, allowing these diatoms to fix atmospheric nitrogen.<ref>{{Cite journal |last1=Schvarcz |first1=Christopher R. |last2=Wilson |first2=Samuel T. |last3=Caffin |first3=Mathieu |last4=Stancheva |first4=Rosalina |last5=Li |first5=Qian |last6=Turk-Kubo |first6=Kendra A. |last7=White |first7=Angelicque E. |last8=Karl |first8=David M. |last9=Zehr |first9=Jonathan P. |last10=Steward |first10=Grieg F. |date=2022-02-10 |title=Overlooked and widespread pennate diatom-diazotroph symbioses in the sea |journal=Nature Communications |language=en |volume=13 |issue=1 |pages=799 |bibcode=2022NatCo..13..799S |doi=10.1038/s41467-022-28065-6 |issn=2041-1723 |pmc=8831587 |pmid=35145076}}</ref><ref>{{Cite journal |pmc=4128115 |year=2014 |last1=Nakayama |first1=T. |title=Complete genome of a nonphotosynthetic cyanobacterium in a diatom reveals recent adaptations to an intracellular lifestyle |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=111 |issue=31 |pages=11407–11412 |last2=Kamikawa |first2=R. |last3=Tanifuji |first3=G. |last4=Kashiyama |first4=Y. |last5=Ohkouchi |first5=N. |last6=Archibald |first6=J. M. |last7=Inagaki |first7=Y. |pmid=25049384 |doi=10.1073/pnas.1405222111 |bibcode=2014PNAS..11111407N |doi-access=free}}</ref> Other diatoms in symbiosis with nitrogen-fixing cyanobacteria are among the genera ''Hemiaulus'', ''Rhizosolenia'' and ''Chaetoceros''.<ref>{{Cite journal |last1=Pierella Karlusich |first1=Juan José |last2=Pelletier |first2=Eric |last3=Lombard |first3=Fabien |last4=Carsique |first4=Madeline |last5=Dvorak |first5=Etienne |last6=Colin |first6=Sébastien |last7=Picheral |first7=Marc |last8=Cornejo-Castillo |first8=Francisco M. |last9=Acinas |first9=Silvia G. |last10=Pepperkok |first10=Rainer |last11=Karsenti |first11=Eric |date=2021-07-06 |title=Global distribution patterns of marine nitrogen-fixers by imaging and molecular methods |journal=Nature Communications |language=en |volume=12 |issue=1 |pages=4160 |doi=10.1038/s41467-021-24299-y |issn=2041-1723 |pmc=8260585 |pmid=34230473 |bibcode=2021NatCo..12.4160P}}</ref>
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