Editing Hyena (section)

===Phylogeny===
The following cladogram illustrates the phylogenetic relationships between extant and extinct hyaenids based on the morphological analysis by Werdelin & Solounias (1991),<ref name="Werdelin1991">{{cite journal|last1=Werdelin|first1=L.|last2=Solounias|first2=Nikos|title=The Hyaenidae: taxonomy, systematics and evolution.|url=http://foreninger.uio.no/ngf/FOS/pdfs/F&S_30.pdf|journal=Fossils and Strata|volume=30|pages=1–104|year=1991|doi=10.18261/8200374815-1991-01 |isbn=8200374815 }}</ref> as updated by Turner et al. (2008).<ref>{{cite journal|title=Taxonomy and evolutionary patterns in the fossil Hyaenidae of Europe|first1=Alan|last1=Turner|first2=Mauricio|last2=Antón|first3=Lars|last3=Werdelin|journal=Geobios|volume=41|issue=5|year=2008|pages=677–687|doi=10.1016/j.geobios.2008.01.001|bibcode=2008Geobi..41..677T }}</ref>
{{Clade
 |style=font-size:80%; line-height:85%;background-color:#eeeeff
 |caption=Evolution of hyaenas
 |captionstyle=background-color:#8899ff;padding:10px;font-weight:bold;
 |footer=Phylogenic relationships based on morphological characteristics, after Werdelin & Solounias (1991) and Turner et al (2008)
 |footerstyle=background-color:#eeeeee;text-align:center;
 |1={{clade
  |style=background-color:#ccffcc;
  |label1=
  |1=''[[Protictitherium|Protictitherium crassum]]''
  |2=''"Protictitherium" cingulatum''
  |3=''"Protictitherium" intermedium''
  |4=''"Protictitherium" llopisi''
  |5=''"Protictitherium" punicum''
  |6=''"[[Protictitherium|Protictitherium" gaillardi]]''
  |state7=dotted
  |7=''"Protictitherium" sumegense''
  |state8=dotted
  |8=''"Protictitherium" csakvarense''
  |style9=background-color:#eeeeff;
  |9={{clade
    |label1=
    |1={{clade
        |label1=''[[Plioviverrops]]''
        |style1=background-color:#ccffcc;
        |1={{clade
           |grouplabel1=Civet/mongoose-like insectivore-omnivores 
           |grouplabelstyle1=font-size:12px;font-weight:bold;color:darkgreen;text-align:right;
           |label1=
           |1=''Plioviverrops gervaisi''
           |2={{clade
              |1=''Plioviverrops orbignyi''
              |2={{Clade
                 |label1=
                 |1=''Plioviverrops guerini''
                 |2=''Plioviverrops faventinus''
                 |3=''Plioviverrops gaudryi''
                  }} 
               }} 
            }}
         }}
     |2={{clade
        |label1=
        |1=''Tungurictis spocki''
        |2={{clade
           |label1=
           |1=''[[Thalassictis|Thalassictis robusta]]''
           |2=''"Thalassictis" certa''
           |3=''"Thalassictis" montadai''
           |4=''"Thalassictis" proava''
           |5=''"Thalassictis" sarmatica''
           |6=''"Thalassictis" spelaea''
           |7={{clade
              |label1=
              |1=''Tongxinictis primordialis''
              |2={{clade
                 |label1=
                 |1={{clade
                    |label1=''[[Proteles]]''
                    |style1=background-color:#ddeeff;
                    |1={{clade
                       |label1=
                       |1='''''[[Proteles cristatus]]''''' (aardwolf) [[File:The life of animals (Colored Plate 4) (proteles cristatus).jpg|50 px]]
                       |2=''Proteles amplidentus''
                        }}
                     }}
                 |2={{clade
                    |label1=
                    |1={{clade
                       |label1=''[[Ictitherium]]''
                       |style=background-color:#ddddff;
                       |1={{clade
                          |label1=
                          |1=''[[Ictitherium|Ictitherium viverrinum]]''
                          |2=''Ictitherium ebu''
                          |grouplabel2=Jackal-like hyaenas 
                          |grouplabelstyle2=font-size:12px;font-weight:bold;color:darkcyan;text-align:right;
                          |3={{clade
                             |label1=
                             |1={{clade
                                |label1=
                                |1=''Ictitherium  tauricum''
                                |2=''Ictitherium ibericum''
                                 }}
                             |2={{clade
                                |label1=
                                |1=''Ictitherium kurteni''
                                |2={{clade
                                   |1=''Ictitherium intuberculatum''
                                   |2=''Ictitherium pannonicum''
                                    }}
                                 }} 
                              }} 
                           }} 
                        }}
                    |2={{clade 
                       |label1=
                       |1=''Miohyaenotherium bessarabicum''
                       |2={{clade
                          |1={{clade
                             |label1=''[[Hyaenotherium]]''
                             |1={{clade
                                |1=''Hyaenotherium wongii''
                                |2=''Hyaenictitherium hyaenoides''
                                |3=''"Hyaenictitherium" pilgrimi''
                                |4=''"Hyaenictitherium" parvum''
                                |5=''"Hyaenictitherium" namaquensis''
                                |6=''"Hyaenictitherium" minimum''
                                 }}
                              }}
                          |style2=background-color:#ccccff;
                          |2={{clade
                             |1={{clade
                                |label1=''[[Lycyaena]]''
                                |1={{clade
                                   |grouplabel2=Cursorial hunting hyaenas 
                                   |grouplabelstyle2=font-size:12px;font-weight:bold;color:darkblue;text-align:right;
                                   |1=''Lycyaena chaeretis''
                                   |2=''Lycyaena dubia''
                                   |3={{clade
                                      |1=''Lycyaena macrostoma''
                                      |2=''Lycyaena crusafonti''
                                       }}
                                    }} 
                                 }}
                             |2={{clade
                                |1={{clade
                                   |label1=''[[Hyaenictis]]''
                                   |1={{clade
                                      |1=''Hyaenictis graeca''
                                      |2=''Hyaenictis almerai''
                                      |3=''Hyaenictis hendeyi''
                                       }}
                                    }}
                                 |2={{clade
                                    |1={{clade
                                       |label1=''[[Lycyaenops]]''
                                       |1={{clade
                                          |1=''Lycyaenops rhomboideae''
                                          |2=''Lycyaenops silberbergi''
                                           }}
                                        }}
                                    |2={{clade
                                       |label1=''[[Chasmaporthetes]]''
                                       |sublabel1=(running hyaenas)
                                       |1={{clade
                                          |1={{clade
                                             |1=''Chasmaporthetes exitelus''
                                             |2=''Chasmaporthetes bonisi''
                                              }}
                                          |2={{clade
                                             |1=''Chasmaporthetes borissiaki''
                                             |2={{clade
                                                |1={{clade
                                                   |1=''Chasmaporthetes lunensis''
                                                   |2=''Chasmaporthetes melei''
                                                    }}
                                                |2={{clade
                                                   |1={{clade
                                                      |1=''Chasmaporthetes ossifragus''
                                                      |2=''Chasmaporthetes'' sp. Florida
                                                       }}
                                                   |2={{clade
                                                      |1=''Chasmaporthetes nitidula''
                                                      |2=''Chasmaporthetes australis''
                                                       }} }} }} }} }}
                                       |style2=background-color:#ddccff
                                       |label2=[[Hyaeninae]]
                                       |sublabel2=(bone-cracking hyenas)
                                       |2={HYAENINAE}
                                         }} }} }} }} }} }} }} }} }} }} }} }} }} 
|targetA={HYAENINAE}
|subcladeA={{clade
  |caption=Bone-cracking hyaenas
  |captionstyle=font-size:12px;font-weight:bold;color:darkred;text-align:right;padding:5px 20px 0px 0px;
  |1=''Metahyaena confector''
  |2=''Palinhyaena reperta''
  |3={{clade
     |1=Hyaenid sp. E Langebaar
     |2={{clade
        |1=''Belbus beaumonti''
    <!--|1=''Leecyaena lycyaenoides''
        |2=''"Leecyaena" bosei'' -->
        |style2=background-color:#eeccff;
        |2={{clade
           |1={{clade
              |1={{clade
                 |1=''Hyaena abronia''
                 |2='''''[[Hyaena hyaena]]''''' (striped hyaena) [[File:Hyaena striata - 1818-1842 - Print - Iconographia Zoologica - Special Collections University of Amsterdam -(white background).jpg|50 px]]
                  }}
              |2={{clade
                 |1=''Parahyaena howelli''
                 |2='''''[[Parahyaena brunnea]]''''' (brown hyaena) [[File:Hyaena fusca (white background).jpg|50 px]]
                  }} }}
           |2={{clade
              |1=''Pliocrocuta perrieri''
              |2={{clade
                 |1=''[[Pachycrocuta|Pachycrocuta brevirostris]]'' (giant hyaena) [[File:Pachycrocuta brevirostris restoration.jpg|70 px]]
             <!--|2=''"Pachycrocuta"'' bellax''-->
                 |2={{clade
                    |1=''[[Adcrocuta|Adcrocuta eximia]]'' [[File:Adcrocuta eximia restoration.jpg|60 px]]
                    |2=''Allohyaena kadici''
                    |3={{clade
                       |1='''''[[Crocuta crocuta]]''''' (spotted hyaena) [[File:Spotted hyena (Crocuta crocuta) by Connor Ashbridge.png|50 px]]
                       |2=''Crocuta eturono''
                   <!--|2=''Crocuta sivalensis''
                       |3=''Crocuta dietrichi'' -->
}} }} }} }} }} }} }} }} }}

A more recent molecular analysis agrees on the phylogenetic relationship between the four extant hyaenidae species (Koepfli ''et al'', 2006<ref name="Koepfli2006">{{cite journal|last1=Koepfli|first1=K.-P.|last2=Jenks|first2=S. M.|last3=Eizirik|first3=E.|last4=Zahirpour|first4=T.|last5=Van Valkenburgh|first5=B.|last6=Wayne|first6=R. K.|title=Molecular systematics of the Hyaenidae: Relationships of a relictual lineage resolved by a molecular supermatrix|journal=Molecular Phylogenetics and Evolution|volume=38|issue=3|date=2006|pages=603–620|doi=10.1016/j.ympev.2005.10.017|pmid=16503281|bibcode=2006MolPE..38..603K }}</ref>).

{{clade|style=font-size:90%; line-height:100%;
|label1='''Hyaenidae'''
|1={{clade
  |label1=
  |1={{clade
     |label1=
     |1=''[[Aardwolf|Proteles cristatus]]'' (aardwolf) [[File:The life of animals (Colored Plate 4) (proteles cristatus).jpg|60 px]]
     }}
  |2={{clade
     |label1=
     |1={{clade |label1=|1=''[[Spotted hyena|Crocuta crocuta]]'' (spotted hyena) [[File:Spotted hyena (Crocuta crocuta) by Connor Ashbridge.png|80 px]] }}
     |2={{clade
        |1=''[[Striped hyena|Hyaena hyaena]]'' (striped hyena) [[File:Hyaena striata - 1818-1842 - Print - Iconographia Zoologica - Special Collections University of Amsterdam -(white background).jpg|70 px]]
        |2=''[[Brown hyena|Parahyaena brunnea]]'' (brown hyena) [[File:Hyaena fusca (white background).jpg|65 px]] 
         }} }} }} 
         
}}