Editing Viroid (section)

===Ambiviruses===
"Ambiviruses" are mobile genetic elements that were recently (2020s) discovered in [[fungi]]. Their RNA [[genome]]s are circular, circa 5 kb in length. One of at least two open reading frames encodes a viral RNA-directed [[RNA polymerase]], that firmly places "ambiviruses" into [[riboviria]]n kingdom ''[[Orthornavirae]]''; a separate phylum ''[[Ambiviricota]]'' has been established since the 2023 ICTV Virus Taxonomy Release because of the unique features of encoding RNA-directed RNA polymerases but also having divergent [[ribozyme]]s in various combinations in both sense and antisense orientation – the detection of circular forms in both sense orientations suggest that "ambiviruses" use rolling circle replication for propagation.<ref>{{cite journal |last1=Sutela |first1=Suvi |last2=Forgia |first2=Marco |last3=Vainio |first3=Eeva J |last4=Chiapello |first4=Marco |last5=Daghino |first5=Stefania |last6=Vallino |first6=Marta |last7=Martino |first7=Elena |last8=Girlanda |first8=Mariangela |last9=Perotto |first9=Silvia |last10=Turina |first10=Massimo |title=The virome from a collection of endomycorrhizal fungi reveals new viral taxa with unprecedented genome organization |journal=Virus Evolution |date=1 July 2020 |volume=6 |issue=2 |pages=veaa076 |doi=10.1093/ve/veaa076 |pmid=33324490 |pmc=7724248}}</ref><ref>{{cite journal |last1=Chong |first1=Li Chuin |last2=Lauber |first2=Chris |title=Viroid-like RNA-dependent RNA polymerase-encoding ambiviruses are abundant in complex fungi |journal=Frontiers in Microbiology |date=12 May 2023 |volume=14 |doi=10.3389/fmicb.2023.1144003 |doi-access=free |pmid=37275138 |pmc=10237039}}</ref><ref name=":2">{{cite web |title=ICTV Virus Taxonomy: 2023 Release |url=https://ictv.global/taxonomy |publisher=ICTV |access-date=2 May 2024}}</ref>