Editing Chronobiology (section)

===Distinct ipRGCs===
Further research has shown that ipRGCs project to different brain nuclei to control both non-image forming and image forming functions.<ref name=":0">{{Cite journal|title = Blurring the boundaries of vision: novel functions of intrinsically photosensitive retinal ganglion cells|last = Matynia|first = Anna|date = September 3, 2013|journal = Journal of Experimental Neuroscience|doi = 10.4137/JEN.S11267|pmc = 4089729|pmid=25157207|volume=7|pages=43–50}}</ref> These brain regions include the SCN, where input from ipRGCs is necessary to photoentrain circadian rhythms, and the [[pretectal area|olivary pretectal nucleus]] (OPN), where input from ipRGCs control the pupillary light reflex.<ref name=":1">{{Cite journal|title = Retinal Ganglion Cell Maps in the Brain: Implications for Visual Processing|last1 = Dhande|first1 = OS|date = November 19, 2013|journal = Current Opinion in Neurobiology|doi = 10.1016/j.conb.2013.08.006|pmc = 4086677|last2 = Huberman|first2 = AD|volume=24|issue = 1|pages=133–142|pmid=24492089}}</ref> Hattar and colleagues conducted research that demonstrated that ipRGCs project to hypothalamic, thalamic, stratal, brainstem and limbic structures.<ref>{{Cite journal|title = Neuroimaging, Cognition, Light and Circadian Rhythms|author1=Gaggioni G |author2=Maquet P |author3=Schmidt C |author4=Dijk Dj |author5=Vandealle G |date = July 8, 2014|journal = Frontiers in Systems Neuroscience|doi = 10.3389/fnsys.2014.00126|pmc = 4086398|pmid=25071478|volume=8|pages=126|doi-access=free }}</ref> Although ipRGCs were initially viewed as a uniform population, further research revealed that there are several subtypes with distinct morphology and physiology.<ref name=":0" /> Since 2011, Hattar's laboratory<ref>{{cite web|title=The Hattar Lab|url=http://hattarlab.johnshopkins.edu/pages/research.php|publisher=Johns Hopkins University|accessdate=27 December 2016|date=2014}}</ref> has contributed to these findings and has successfully distinguished subtypes of ipRGCs.<ref name=":1"/>