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===Recognizing variation=== In the late 1980s and early 1990s, S. Christopher Bennett published several major papers reviewing the anatomy, taxonomy and life history of ''Pteranodon''.<ref name= kellner2010/> In 1992, he published a paper discussing [[sexual dimorphism]] and its role in individual variation among ''Pteranodon'' fossils, a follow-up of a 1987 paper he authored on the same subject.<ref name=":1">{{Cite journal |last=Bennett |first=S. Christopher |date=1987 |title=Sexual dimorphism in the pterosaur Pteranodon. |journal=Journal of Vertebrate Paleontology |volume=7}}</ref> In the 1992 paper, he referred only to two species, ''P. longiceps'' and ''P. sternbergi''.<ref name="bennett1992" /> Two years later, he published a paper fully revising its taxonomy, wherein he concluded that only ''P. longiceps'' and ''P. sternbergi'' were valid species. ''P. marshi'' and ''P. walkeri'' were regarded as junior synonyms of ''P. longiceps'', and ''P. eatoni'' as a junior synonym of ''P. stenbergi''. The remainder were either rendered ''nomina dubia'' or placed in Nyctosaurus''.''<ref name="bennett1994" />
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