Editing Maotianshan Shales (section)

== Guanshan biota ==
{{further|List of Guanshan Biota species}}
Located in the Yunnan Province of South China and hosted in the geologically distinct [[Cambrian Stage 4]] [[Wulongqing Formation]], The Guanshan biota are also Burgess shale-type fossils but slightly younger than the Chengjian biota  with an age dating to 515–510 [[Myr]].<ref>{{Cite journal |last1=Chen |first1=Feiyang |last2=Zhang |first2=Zhifei |last3=Betts |first3=Marissa J. |last4=Zhang |first4=Zhiliang |last5=Liu |first5=Fan |date=2019 |title=First report on Guanshan Biota (Cambrian Stage 4) at the stratotype area of Wulongqing Formation in Malong County, Eastern Yunnan, China |journal=Geoscience Frontiers |language=en |volume=10 |issue=4 |pages=1459–1476 |doi=10.1016/j.gsf.2018.09.010|bibcode=2019GeoFr..10.1459C |s2cid=134921385 |doi-access=free }}</ref><ref name=":0">{{Cite journal |last1=Zhao |first1=Jun |last2=Li |first2=Yujing |last3=Selden |first3=Paul A. |last4=Cong |first4=Peiyun |date=2020 |title=New occurrence of the Guanshan Lagerstätte (Cambrian Series 2, Stage 4) in the Kunming area, Yunnan, southwest China, with records of new taxa |url=https://www.tandfonline.com/doi/full/10.1080/03115518.2020.1781257 |journal=Alcheringa: An Australasian Journal of Palaeontology |language=en |volume=44 |issue=3 |pages=343–355 |doi=10.1080/03115518.2020.1781257 |bibcode=2020Alch...44..343Z |s2cid=221093834 |issn=0311-5518 |access-date=2023-03-18 |archive-date=2023-03-18 |archive-url=https://web.archive.org/web/20230318052728/https://www.tandfonline.com/doi/full/10.1080/03115518.2020.1781257 |url-status=live }}</ref> Brachiopods are the most abundant species,<ref>{{Cite journal |last1=Hu |first1=Shixue |last2=Zhang |first2=Zhifei |last3=Holmer |first3=Lars E. |last4=Skovsted |first4=Christian B. |date=2010 |title=Soft-Part Preservation in a Linguliform Brachiopod from the Lower Cambrian Wulongqing Formation (Guanshan Fauna) of Yunnan, South China |url=http://www.app.pan.pl/article/item/app20091106.html |journal=Acta Palaeontologica Polonica |language=en |volume=55 |issue=3 |pages=495–505 |doi=10.4202/app.2009.1106 |s2cid=59439966 |issn=0567-7920 |access-date=2023-03-18 |archive-date=2023-03-18 |archive-url=https://web.archive.org/web/20230318052734/https://www.app.pan.pl/article/item/app20091106.html |url-status=live }}</ref> followed by trilobites. Other species belong to sponges, chancelloriids, cnidarians, ctenophores, priapulids, lobopodians, arthropods, anomalocaridids, hyoliths, molluscs, brachiopods, echinoderms, algae and vetulicolians. There are also the earliest-known eocrinoids, unidentified soft-bodied animals and abundant trace fossils.<ref name=":1">{{Cite journal |last1=Hu |first1=ShiXue |last2=Zhu |first2=MaoYan |last3=Steiner |first3=Michael |last4=Luo |first4=HuiLin |last5=Zhao |first5=FangChen |last6=Liu |first6=Qi |date=2010 |title=Biodiversity and taphonomy of the Early Cambrian Guanshan biota, eastern Yunnan |url=http://link.springer.com/10.1007/s11430-010-4086-9 |journal=Science China Earth Sciences |language=en |volume=53 |issue=12 |pages=1765–1773 |doi=10.1007/s11430-010-4086-9 |bibcode=2010ScChD..53.1765H |s2cid=128882075 |issn=1674-7313}}</ref>

The Guanshan biota are regarded as successors of the Chengjian biota,<ref name=":0" /> and share many species. The unique species include arthropods like ''[[Guangweicaris]]''<ref>{{Cite journal |last1=Wu |first1=Yichen |last2=Liu |first2=Jianni |date=2019 |title=Anatomy and relationships of the fuxianhuiid euarthropod Guangweicaris from the early Cambrian Guanshan Biota in Kunming, Yunnan, Southwest China revisited |url=http://www.app.pan.pl/article/item/app005422018.html |journal=Acta Palaeontologica Polonica |volume=64 |doi=10.4202/app.00542.2018 |s2cid=201291723 |doi-access=free |access-date=2023-03-18 |archive-date=2023-03-18 |archive-url=https://web.archive.org/web/20230318054121/https://www.app.pan.pl/article/item/app005422018.html |url-status=live }}</ref> and ''[[Astutuscaris]],''<ref>{{Cite journal |last1=Jiao |first1=De guang |last2=Du |first2=Kunsheng |date=2022 |title=A new euarthropod from the Cambrian Stage 4 Guanshan Biota of South China |url=https://www.app.pan.pl/article/item/app009372021.html |journal=Acta Palaeontologica Polonica |volume=67 |doi=10.4202/app.00937.2021 |s2cid=253068676 |doi-access=free |access-date=2023-03-18 |archive-date=2023-03-18 |archive-url=https://web.archive.org/web/20230318054119/https://www.app.pan.pl/article/item/app009372021.html |url-status=live }}</ref>alongside vetulicolians like ''[[Vetulicola gantoucunensis]]''<ref>{{Cite journal |last1=Huilin |first1=Luo |last2=Xiaoping |first2=Fu |last3=Shixue |first3=Hu |last4=Yong |first4=Li |last5=Liangzhong |first5=Chen |last6=Ting |first6=You |last7=Qi |first7=Liu |date=2005 |title=New Vetulicoliids from the Lower Cambrian Guanshan Fauna, Kunming |url=https://onlinelibrary.wiley.com/doi/10.1111/j.1755-6724.2005.tb00860.x |journal=Acta Geologica Sinica - English Edition |language=en |volume=79 |issue=1 |pages=1–6 |doi=10.1111/j.1755-6724.2005.tb00860.x |bibcode=2005AcGlS..79....1L |s2cid=129031316 |access-date=2023-03-18 |archive-date=2023-03-08 |archive-url=https://web.archive.org/web/20230308053427/https://onlinelibrary.wiley.com/doi/10.1111/j.1755-6724.2005.tb00860.x |url-status=live }}</ref> and ''[[Vetulicola longbaoshanensis|V. longbaoshanensis]]''.<ref>{{Cite journal |last1=Li |first1=JinShu |last2=Liu |first2=JianNi |last3=Ou |first3=Qiang |date=2017 |title=New observations on Vetulicola longbaoshanensis from the Lower Cambrian Guanshan Biota (Series 2, Stage 4), South China |url=http://link.springer.com/10.1007/s11430-017-9088-y |journal=Science China Earth Sciences |language=en |volume=60 |issue=10 |pages=1795–1804 |doi=10.1007/s11430-017-9088-y |bibcode=2017ScChD..60.1795L |s2cid=135037211 |issn=1674-7313}}</ref>