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=== Trunk and lobopods === {{multiple image | align = right | width = 220 | footer = | image1 = 20210829 Paucipodia inermis diagrammatic reconstruction.png | caption1 = ''[[Paucipodia]] inermis'', a lobopodian with featureless, undifferentiated trunk region. | image2 = 20210902 Luolishania longicruris Miraluolishania haikouensis diagrammatic reconstruction.png | caption2 = ''[[Luolishania]] longicruris'', showing clear differentiation between trunk segments and lobopods. }} The trunk is elongated and composed of numerous body segments ([[somites]]), each bearing a pair of legs called lobopods<ref name=":0" /> or lobopodous limbs.<ref name=":1" /> The segmental boundaries are not as externally significant as those of arthropods, although they are indicated by heteronomous annulations (i.e., the alternation of annulation density corresponding to the position of segmental boundaries) in some species.<ref name=":33">Chen, J.Y., Zhou, G.Q., Ramsköld, L. (1995a). [https://www.academia.edu/3776072 The Cambrian lobopodian ''Microdictyon sinicum'']. Bulletin of the National Museum of Natural Science 5, 1–93 (Taichung, Taiwan).</ref><ref name=":4" /><ref name="ReferenceA" /> The trunk segments may bear other external, segment-corresponding structures such as nodes (e.g. ''[[Hadranax]]'',<ref name=":16" /> ''[[Kerygmachela]]''<ref name=":1" />), papillae (e.g. ''[[Onychodictyon]]''<ref name=":22" />), spine/plate-like sclerites (e.g. armoured lobopodians<ref name=":0" />) or lateral flaps (e.g. gilled lobopodians<ref name=":1" /><ref name=":7" />). The trunk may terminate with a pair of lobopods (e.g. ''[[Aysheaia]]'', ''[[Hallucigenia]] sparsa'')<ref name=":2" /> or a tail-like extension (e.g. ''[[Paucipodia]]'', ''[[Siberion]]'', ''[[Jianshanopodia]]'').<ref name=":33" /><ref name="Hou2004" /><ref name="Liu2006" /><ref name=":9" /> The lobopods are flexible and loosely conical in shape, tapering from the body to tips that may <ref name="ReferenceA" /><ref name=":0" /> or may not <ref name=":3">{{cite journal |last1=Ou |first1=Qiang |last2=Liu |first2=Jianni |last3=SHU |first3=DEGAN |last4=Han |first4=Jian |last5=Zhang |first5=Zhifei |last6=Wan |first6=Xiaoqiao |last7=Lei |first7=Qianping |date=2011-05-01 |title=A Rare Onychophoran-Like Lobopodian from the Lower Cambrian Chengjiang Lagerstätte, Southwestern China, and its Phylogenetic Implications |url=https://www.researchgate.net/publication/259887599 |journal=Journal of Paleontology |volume=85 |issue=3 |pages=587–594 |doi=10.1666/09-147R2.1 |jstor=23020193|bibcode=2011JPal...85..587O |s2cid=53056128}}</ref><ref name=":12">{{cite journal|last1=Ma|first1=Xiaoya|last2=Edgecombe|first2=Gregory|last3=Legg|first3=David|last4=Hou|first4=Xianguang|date=2013-05-08|title=The morphology and phylogenetic position of the Cambrian lobopodian ''Diania cactiformis''|url=https://www.researchgate.net/publication/247777382|journal=Journal of Systematic Palaeontology|volume=12|issue=4|pages=445–457|doi=10.1080/14772019.2013.770418|s2cid=220463025}}</ref><ref name=":13">{{cite journal|last1=Ou|first1=Qiang|last2=Mayer|first2=Georg|date=2018-12-01|title=A Cambrian unarmoured lobopodian, †''Lenisambulatrix humboldti'' gen. et sp. nov., compared with new material of †''Diania cactiformis''|url=https://www.researchgate.net/publication/327777757|journal=Scientific Reports|volume=8|issue=1|pages=13667|bibcode=2018NatSR...813667O|doi=10.1038/s41598-018-31499-y|pmc=6147921|pmid=30237414|doi-access=free}}</ref> bear claws. The claws, if present, are hardened structures with a shape resembling a hook or gently-curved spine.<ref name="Hou2004" /><ref name=":36">{{Cite journal|last1=Steiner|first1=M.|last2=Hu|first2=S.X.|last3=Liu|first3=J.|last4=Keupp|first4=H.|date=2012-02-02|title=A new species of ''Hallucigenia'' from the Cambrian Stage 4 Wulongqing Formation of Yunnan (South China) and the structure of sclerites in lobopodians|journal=Bulletin of Geosciences|pages=107–124|doi=10.3140/bull.geosci.1280|issn=1802-8225|doi-access=free}}</ref><ref name=":4" /><ref name="ReferenceA" /><ref name=":0" /> Claw-bearing lobopods usually have two claws, but single claws are known (e.g. posterior lobopods of [[luolishaniids]]<ref name=":4" /><ref name=":6" /><ref name=":28" />), as are more than two (e.g. three in ''[[Tritonychus]]'',<ref name=":34" /> seven in ''[[Aysheaia]]''<ref name=":32" />) depending on its segmental or taxonomical association.<ref name="ReferenceA" /> In some genera, the lobopods bear additional structures such as spines (e.g. ''[[Diania]]''<ref name=":12" />), fleshy outgrowths (e.g. ''[[Onychodictyon]]''<ref name=":22" />), or tubercules (e.g. ''[[Jianshanopodia]]''<ref name="Liu2006" />). There is no sign of [[Arthropod leg|arthropodization]] (development of a hardened exoskeleton and segmental division on panarthropod appendages) in known members of lobopodians, even for those belonging to the [[arthropod]] stem-group (e.g. gilled lobopodians and siberiids), and the suspected case of arthropodization on the limbs of ''[[Diania]]''<ref name=":24">{{cite journal |last1=Liu |first1=Jianni |last2=Steiner |first2=Michael |last3=Dunlop |first3=Jason A. |last4=Keupp |first4=Helmut |last5=Shu |first5=Degan |last6=Ou |first6=Qiang |last7=Han |first7=Jian |last8=Zhang |first8=Zhifei |last9=Zhang |first9=Xingliang |date=February 2011 |title=An armoured Cambrian lobopodian from China with arthropod-like appendages |journal=Nature |language=en |volume=470 |issue=7335 |pages=526–530 |doi=10.1038/nature09704 |pmid=21350485 |issn=1476-4687 |bibcode=2011Natur.470..526L|s2cid=4324509}}</ref> is considered to be a misinterpretation.<ref name=":12" /><ref name=":13" /> Differentiation (tagmosis) between trunk somites barely occurs, except in [[hallucigenids]] and luolishaniids, where numerous pairs of their anterior lobopods are significantly slender (hallucigenids) or setose (luolishaniids) in contrast to their posterior counterparts.<ref name=":0" /><ref name=":2" /><ref name=":6" /><ref name=":11" /><ref name=":28" />
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