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==Fossil record== [[File:Hydrangea knowltoni as Hydrangea bendirei USNM P36979 img3.jpg|thumb|right|''[[Hydrangea knowltoni]]'']] {{extinct}}''[[Hydrangea alaskana]]'' is a fossil species recovered from [[Paleogene]] strata at Jaw Mountain Alaska.<ref>{{cite journal |last=Hollick |first=Arthur |year=1925 |title=A New Fossil Species of Hydrangea |journal=Bulletin of the Torrey Botanical Club |volume=52 |issue=1 |pages=21–22 |doi=10.2307/2479996 |jstor=2479996}}</ref> †''[[Hydrangea knowltoni]]'' has been described from leaves and flowers recovered from the Miocene [[Langhian]] [[Latah Formation]] of the inland Pacific Northwest United states. The related Miocene species †''[[Hydrangea bendirei]]'' is known to from the [[Mascall Formation]] in Oregon, and †''[[Hydrangea reticulata]]'' is documented from the [[Weaverville Formation]] in California.<ref name="Knowlton1926">{{cite report |last1=Knowlton |first1=F.H. |year=1926 |chapter=Flora of the Latah Formation of Spokane, Washington, and Coeur d'Alene, Idaho |title=Shorter contributions to general geology, 1925 |series=Professional Paper |publisher=United States Geological Survey |volume=140 |pages=17–55, plates VIII-XXXI |doi=10.3133/pp140A |url=https://pubs.usgs.gov/pp/0140a/report.pdf}}</ref><ref name="Columbia1959">{{cite book |last1=Chaney |first1=R. |last2=Axelrod |first2=D. |year=1959 |title=Miocene Floras of the Columbia Plateau: Part II. Systematic Considerations, by Ralph W. Chaney and Daniel I. Axelrod |publisher= Carnegie Institution of Washington |pages=1–226}}{{HathiTrust Catalog|uc1.c099384505|Miocene Floras of the Columbia Plateau}}</ref> Four [[fossil]] seeds of †''[[Hydrangea polonica]]'' have been extracted from [[borehole]] samples of the [[Middle Miocene]] fresh water deposits in [[Nowy Sacz]] Basin, [[West Carpathians]], [[Poland]].<ref>Łańcucka-Środoniowa M.: Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) [Szczątki makroskopowe roślin z miocenu słodkowodnego Kotliny Sądeckiej (Karpaty Zachodnie, Polska)]. Acta Palaeobotanica 1979 20 (1): 3-117.</ref>
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