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=== Invertebrates === {{further|Respiratory system of gastropods}} [[File:Spider internal anatomy-en.svg|thumb|[[Book lung]]s of a female spider (shown in pink)]] A number of [[invertebrate]]s have lung-like structures that serve a similar respiratory purpose to true vertebrate lungs, but are not evolutionarily related and only arise out of [[convergent evolution]]. Some [[arachnid]]s, such as [[spider]]s and [[scorpion]]s, have structures called [[book lung]]s used for atmospheric gas exchange. Some species of spider have four pairs of book lungs but most have two pairs.<ref>{{Cite web | url = https://www.britannica.com/science/book-lung | title = book lung {{!}} anatomy | website = Encyclopædia Britannica | access-date = 2016-02-24 }}</ref> Scorpions have [[Spiracle (arthropods)|spiracle]]s on their body for the entrance of air to the book lungs.<ref>{{Cite web | url = https://www.britannica.com/science/spiracle | title = spiracle {{!}} anatomy | website = Encyclopædia Britannica | access-date = 2016-02-24 }}</ref> The [[coconut crab]] is terrestrial and uses structures called [[branchiostegal lung]]s to breathe air.<ref name="Farrelly2005">{{cite journal|vauthors=Farrelly CA, Greenaway P|year=2005|title=The morphology and vasculature of the respiratory organs of terrestrial hermit crabs (''Coenobita'' and ''Birgus''): gills, branchiostegal lungs and abdominal lungs |journal=Arthropod Structure & Development|volume=34 |issue=1 |pages=63–87 |doi=10.1016/j.asd.2004.11.002|bibcode=2005ArtSD..34...63F }}</ref> Juveniles are released into the ocean, however adults cannot swim and possess an only rudimentary set of gills. The adult crabs can breathe on land and hold their breath underwater.<ref>{{Cite book | url = https://books.google.com/books?id=RR09AAAAIAAJ | title = Biology of the Land Crabs | last1 = Burggren | first1 = Warren W. | last2 = McMahon | first2 = Brian R. | year = 1988 | publisher = Cambridge University Press | isbn = 978-0-521-30690-4 | page = 25 | language = en }}</ref> The branchiostegal lungs are seen as a developmental adaptive stage from water-living to enable land-living, or from fish to amphibian.<ref>{{Cite book | url = https://books.google.com/books?id=RR09AAAAIAAJ | title = Biology of the Land Crabs | last1 = Burggren | first1 = Warren W. | last2 = McMahon | first2 = Brian R. | year = 1988 | publisher = Cambridge University Press | isbn = 978-0-521-30690-4 | page = 331 | language = en }}</ref> [[Pulmonates]] are mostly [[land snail]]s and [[slug]]s that have developed a simple lung from the [[mantle cavity]]. An externally located opening called the [[pneumostome]] allows air to be taken into the mantle cavity lung.<ref>Land Snails (& other Air-Breathers in Pulmonata Subclass & Sorbeconcha Clade). at Washington State University Tri-Cities Natural History Museum. Accessed 25 February 2016. http://shells.tricity.wsu.edu/ArcherdShellCollection/Gastropoda/Pulmonates.html {{Webarchive|url=https://web.archive.org/web/20181109010506/http://shells.tricity.wsu.edu/ArcherdShellCollection/Gastropoda/Pulmonates.html |date=2018-11-09 }}</ref><ref>{{Cite book | url = https://books.google.com/books?id=7UyeBQAAQBAJ | title = Mollusca: Metabolic Biochemistry and Molecular Biomechanics | last = Hochachka | first = Peter W. | year = 2014 | publisher = Academic Press | isbn = 978-1-4832-7603-8 | language = en }}</ref>
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