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=== Evolution === The fossilized wing of an indeterminate locust has been found in [[Rupelian|Early Oligocene]]-aged sediments of the [[Pabdeh Formation]] in [[Iran]], which were deposited in a deep marine environment. The locust was likely migrating across the early [[Paratethys|Paratethys Sea]], between the emergent Arabian Peninsula and central Iran, which were still separated by large areas of deep ocean at this time. This suggests that trans-oceanic locust migrations have been occurring for at least 30 million years, likely facilitated by the spread of [[Grassland|grasslands]] at the time.<ref>{{Cite journal |last1=Mirzaie Ataabadi |first1=Majid |last2=Bahrami |first2=Ali |last3=Yazdi |first3=Mehdi |last4=Nel |first4=AndrΓ© |date=2019-05-28 |title=A locust witness of a trans-oceanic Oligocene migration between Arabia and Iran (Orthoptera: Acrididae) |url=https://www.tandfonline.com/doi/full/10.1080/08912963.2017.1378651 |journal=Historical Biology |language=en |volume=31 |issue=5 |pages=574β580 |doi=10.1080/08912963.2017.1378651 |bibcode=2019HBio...31..574M |issn=0891-2963}}</ref>
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