Editing Cat (section)

=== Domestication ===
{{See also|Domestication of the cat|Cats in ancient Egypt}}
[[File:Tomb of Nakht (7).jpg|thumb|A cat eats a fish under a chair, in a [[mural]] in an Egyptian tomb dating to the 15th century BC.]]
It was long thought that the [[domestication of the cat]] began in [[ancient Egypt]], where cats were venerated from around 3100 BC.<ref>{{cite book |title=The Cat in ancient Egypt, illustrated from the collection of cat and other Egyptian figures formed |publisher=Cambridge University Press |last1=Langton |first1=N. |last2=Langton |first2=M. B. |date=1940}}</ref><ref>{{cite book |title=The Cat in Ancient Egypt |publisher=[[University of Pennsylvania Press]] |last=Malek |first=J. |date=1997 |edition=Revised |location=Philadelphia}}</ref> However, the earliest known indication for the [[taming]] of an African wildcat was [[Excavation (archaeology)|excavated]] close by a human [[Neolithic]] grave in [[Shillourokambos]], southern [[Cyprus]], dating to about 7500–7200 BC. Since there is no evidence of native mammalian [[fauna]] on Cyprus, the inhabitants of this Neolithic village most likely brought the cat and other wild mammals to the island from the [[Middle East]]ern mainland.<ref name="Vigne_al2004">{{cite journal |last1=Vigne |first1=J. D. |last2=Guilaine |first2=J. |last3=Debue |first3=K. |last4=Haye |first4=L. |last5=Gérard |first5=P. |s2cid=28294367 |title=Early taming of the cat in Cyprus |journal=[[Science (journal)|Science]] |volume=304 |issue=5668 |page=259 |year=2004 |pmid=15073370 |doi=10.1126/science.1095335}}</ref> Scientists therefore assume that African wildcats were attracted to early human settlements in the [[Fertile Crescent]] by rodents, in particular the [[house mouse]] (''Mus musculus''), and were tamed by Neolithic farmers. This [[Mutualism (biology)|mutual]] relationship between early farmers and tamed cats lasted thousands of years. As [[agricultural practices]] spread, so did tame and domesticated cats.<ref name="DriscollMenotti-Raymond2007">{{Cite journal |title=The Near Eastern Origin of Cat Domestication |last1=Driscoll |first1=C. A. |last2=Menotti-Raymond |first2=M. |last3=Roca '|first3=A. L. |last4=Hupe |first4=K. |last5=Johnson |first5=W. E. |last6=Geffen |first6=E. |last7=Harley |first7=E. H. |last8=Delibes |first8=M. |last9=Pontier |first9=D. |last10=Kitchener |first10=A. C. |last11=Yamaguchi |first11=N. |last12=O'Brien |first12=S. J. |last13=Macdonald |first13=D. W. |journal=[[Science (journal)|Science]] |year=2007 |volume=317 |issue=5837 |pages=519–523 |pmid=17600185 |pmc=5612713 |doi=10.1126/science.1139518 |oclc=808298830 |bibcode=2007Sci...317..519D}}</ref><ref name="Driscoll_al2009">{{cite journal |last1=Driscoll |first1=C. A. |last2=Clutton-Brock |first2=J. |last3=Kitchener |first3=A. C. |last4=O'Brien |first4=S. J. |year=2009 |title=The taming of the cat |journal=Scientific American |volume=300 |issue=6 |pages=68–75 |doi=10.1038/scientificamerican0609-68 |doi-broken-date=1 November 2024 |pmid=19485091 |pmc=5790555 |bibcode=2009SciAm.300f..68D |jstor=26001382}}</ref> Wildcats of Egypt contributed to the maternal [[gene pool]] of the domestic cat at a later time.<ref name="OttoniVan Neer2017">{{cite journal |last1=Ottoni |first1=C. |last2=van Neer |first2=W. |last3=de Cupere |first3=B. |last4=Daligault |first4=J. |last5=Guimaraes |first5=S. |last6=Peters |first6=J. |last7=Spassov |first7=N. |last8=Prendergast |first8=M. E. |last9=Boivin |first9=N. |last10=Morales-Muñiz |first10=A. |last11=Bălăşescu |first11=A. |last12=Becker |first12=C. |last13=Benecke |first13=N. |last14=Boroneant |first14=A. |last15=Buitenhuis |first15=H. |last16=Chahoud |first16=J. |last17=Crowther |first17=A. |last18=Llorente |first18=L. |last19=Manaseryan |first19=N. |last20=Monchot |first20=H. |last21=Onar |first21=V. |last22=Osypińska |first22=M. |last23=Putelat |first23=O. |last24=Quintana Morales |first24=E.M. |last25=Studer |first25=J. |last26=Wierer |first26=U. |last27=Decorte |first27=R. |last28=Grange |first28=T. |last29=Geigl |first29=E. |s2cid=44041769 |title=The palaeogenetics of cat dispersal in the ancient world |journal=Nature Ecology & Evolution |volume=1 |issue=7 |year=2017 |page=0139 |doi=10.1038/s41559-017-0139 |bibcode=2017NatEE...1..139O |url= https://research.rug.nl/en/publications/the-paleogenetics-of-cat-dispersal-in-the-ancient-world(04942e78-fa48-4700-ad97-29fcdf9077a1).html |access-date=18 October 2021 |archive-date=7 March 2022 |archive-url= https://web.archive.org/web/20220307214831/https://research.rug.nl/en/publications/the-paleogenetics-of-cat-dispersal-in-the-ancient-world |url-status=live}}</ref>

The earliest known evidence for the occurrence of the domestic cat in [[Greece]] dates to around 1200 BC. Greek, [[Phoenicia]]n, [[Carthaginia]]n and [[Etrusca]]n traders introduced domestic cats to southern Europe.<ref name=Faure2009>{{cite journal |last1=Faure |first1=E. |last2=Kitchener |first2=A. C. |year=2009 |title=An archaeological and historical review of the relationships between Felids and people |journal=Anthrozoös |volume=22 |issue=3 |page=221−238 |doi=10.2752/175303709X457577|s2cid=84308532}}</ref> By the 5th century BC, they were familiar animals around settlements in [[Magna Graecia]] and [[Etruria]].<ref>{{Cite journal |last1=Ragni |first1=B. |last2=Possenti |first2=M. |last3=Sforzi |first3=A. |last4=Zavalloni |first4=D. |last5=Ciani |first5=F. |year=1994 |title=The wildcat in central-northern Italian peninsula: a biogeographical dilemma |journal=Biogeographia |volume=17 |issue=1 |doi=10.21426/B617110417 |url= https://cloudfront.escholarship.org/dist/prd/content/qt1dz6x5xf/qt1dz6x5xf.pdf |doi-access=free |access-date=29 August 2019 |archive-date=26 July 2018 |archive-url= https://web.archive.org/web/20180726121432/https://cloudfront.escholarship.org/dist/prd/content/qt1dz6x5xf/qt1dz6x5xf.pdf |url-status=live}}</ref> During the [[Roman Empire]], they were introduced to [[Corsica]] and [[Sardinia]] before the beginning of the 1st century AD.<ref name=Vigne1992>{{cite journal |last1=Vigne |first1=J.-D. |title=Zooarchaeology and the biogeographical history of the mammals of Corsica and Sardinia since the last ice age |year=1992 |journal=Mammal Review |volume=22 |issue=2 |pages=87–96 |doi=10.1111/j.1365-2907.1992.tb00124.x|bibcode=1992MamRv..22...87V }}</ref> By the end of the Western Roman Empire in the 5th century, the Egyptian domestic cat lineage had arrived in a [[Baltic Sea]] port in northern [[Germany]].<ref name="OttoniVan Neer2017" />

The [[leopard cat]] (''Prionailurus bengalensis'') was [[tamed]] independently in China around 5500 BC. This line of partially domesticated cats leaves no trace in the domestic cat populations of today.<ref>{{cite journal |title=Earliest 'domestic' cats in China identified as leopard cat (''Prionailurus bengalensis'') |journal=PLOS ONE |year=2016 |page=e0147295 |volume=11 |issue=1 |first1=J.-D. |last1=Vigne |first2=A. |last2=Evin |first3=T. |last3=Cucchi |first4=L. |last4=Dai |first5=C. |last5=Yu |first6=S. |last6=Hu |first7=N. |last7=Soulages |first8=W. |last8=Wang |first9=Z. |last9=Sun |pmid=26799955 |pmc=4723238 |bibcode=2016PLoSO..1147295V |doi=10.1371/journal.pone.0147295 |doi-access=free}}</ref>

During domestication, cats have undergone only minor changes in anatomy and behavior, and they are still capable of surviving in the wild. Several natural behaviors and characteristics of wildcats may have [[Preadaptation|pre-adapted]] them for domestication as pets. These traits include their small size, social nature, obvious body language, love of play, and high intelligence. Their rigorous grooming habits and instinct to bury their bodily waste make them generally much less messy than other domesticated animals. Captive ''[[Leopardus]]'' cats may also display affectionate behavior toward humans but are not domesticated.<ref name=CameronBeaumont>{{cite journal |title=Evidence suggesting pre-adaptation to domestication throughout the small Felidae |last1=Cameron-Beaumont |first1=C. |last2=Lowe |first2=S. E. |last3=Bradshaw |first3=J. W. S. |journal=Biological Journal of the Linnean Society |year=2002 |volume=75 |issue=3 |pages=361–366 |doi=10.1046/j.1095-8312.2002.00028.x |url= https://www.gwern.net/docs/catnip/2002-cameronbeaumont.pdf |doi-access=free |access-date=10 October 2019 |archive-date=10 October 2019 |archive-url= https://web.archive.org/web/20191010072239/https://www.gwern.net/docs/catnip/2002-cameronbeaumont.pdf |url-status=live}}</ref> House cats may mate with feral cats.<ref name=Bradshaw1999>{{cite journal |title=Feral cats: Their role in the population dynamics of ''Felis catus'' |last1=Bradshaw |first1=J. W. S. |last2=Horsfield |first2=G. F. |last3=Allen |first3=J. A. |last4=Robinson |first4=I. H. |journal=Applied Animal Behaviour Science |year=1999 |volume=65 |issue=3 |pages=273–283 |doi=10.1016/S0168-1591(99)00086-6 |url=https://www.gwern.net/docs/catnip/1999-bradshaw.pdf |archive-url= https://web.archive.org/web/20190130202509/https://www.gwern.net/docs/catnip/1999-bradshaw.pdf |url-status=dead |archive-date=30 January 2019}}</ref> [[Felid hybrid#Domestic cat and hybridization|Hybridization between domestic and other Felinae species]] is also possible, producing hybrids such as the [[Kellas cat]] in [[Scotland]].<ref name=Oliveira/><ref>{{cite journal |last1=Kitchener |first1=C. |last2=Easterbee |first2=N. |year=1992 |title=The taxonomic status of black wild felids in Scotland |journal=Journal of Zoology |volume=227 |issue=2 |pages=342–346 |doi=10.1111/j.1469-7998.1992.tb04832.x}}</ref>

Development of [[cat breeds]] started in the mid 19th century.<ref>{{cite book |last=Wastlhuber |first=J. |date=1991 |chapter=History of domestic cats and cat breeds |pages=1–59 |title=Feline Husbandry: Diseases and management in the multiple-cat environment |editor1-last=Pedersen |editor1-first=N. C. |publisher=American Veterinary Publications |location=Goleta |isbn=9780939674299}}</ref> An analysis of the domestic cat [[genome]] revealed that the ancestral wildcat genome was significantly altered in the process of domestication, as specific [[mutation]]s were selected to develop cat breeds.<ref>{{cite journal |last1=Montague |first1=M. J. |last2=Li |first2=G. |last3=Gandolfi |first3=B. |last4=Khan |first4=R. |last5=Aken |first5=B. L. |last6=Searle |first6=S. M. |last7=Minx |first7=P. |last8=Hillier |first8=L. W. |last9=Koboldt |first9=D. C. |last10=Davis |first10=B. W. |last11=Driscoll |first11=C. A. |year=2014 |title=Comparative analysis of the domestic cat genome reveals genetic signatures underlying feline biology and domestication |journal=Proceedings of the National Academy of Sciences |volume=111 |issue=48 |pages=17230–17235 |doi=10.1073/pnas.1410083111 |pmid=25385592 |pmc=4260561 |doi-access=free |bibcode=2014PNAS..11117230M}}</ref> Most breeds are founded on random-bred domestic cats. [[Genetic diversity]] of these breeds varies between regions, and is lowest in purebred populations, which show more than 20 deleterious [[genetic disorder]]s.<ref name=Lipinski>{{cite journal |title=The ascent of cat breeds: Genetic evaluations of breeds and worldwide random-bred populations |last1=Lipinski |first1=M.J. |last2=Froenicke |first2=L. |last3=Baysac |first3=K. C. |last4=Billings |first4=N. C. |last5=Leutenegger |first5=C. M. |last6=Levy |first6=A. M. |last7=Longeri |first7=M. |last8=Niini |first8=T. |last9=Ozpinar |first9=H. |last10=Slater |first10=M.R. |last11=Pedersen |first11=N. C. |last12=Lyons |first12=L. A. |journal=Genomics |year=2008 |volume=91 |issue=1 |pages=12–21 |doi=10.1016/j.ygeno.2007.10.009 |pmid=18060738 |pmc=2267438}}</ref>