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===Ecology=== {{see also|Biodiversity of Borneo|Fauna of Borneo|Flora of Borneo|List of endemic birds of Borneo|List of mammals of Borneo}} [[File:Bornean orangutan (Pongo pygmaeus), Tanjung Putting National Park 01.jpg|thumb|left|The [[critically endangered]] [[Bornean orangutan]], a great ape [[endemic]] to Borneo]] The Borneo [[rainforest]] is estimated to be around 140 million years old, making it one of the oldest rainforests in the world.<ref>{{cite book|author1=Andy Leeder|author2=Alan Brown|author3=Gregg Coleman|author4=Bob Digby|author5=Glyn Owen|author6=Val Davis|title=WJEC GCSE Geography|url=https://books.google.com/books?id=rDjEDAAAQBAJ&pg=PA199-IA444|date=22 August 2016|publisher=Hodder Education|isbn=978-1-4718-6130-7|pages=199β}}</ref> The current dominant tree group, the [[Dipterocarpaceae|dipterocarps]], has dominated the [[Borneo lowland rain forests]] for millions of years.<ref>{{cite news |title=A prehistoric forest grows in Brunei |url=https://www.sciencedaily.com/releases/2022/04/220428104015.htm |work=ScienceDaily |date=28 April 2022}}</ref> It is the centre of the evolution and distribution of many [[Endemism|endemic]] species of plants and animals, and the rainforest is one of the few remaining [[Habitat (ecology)|natural habitats]] for the endangered [[Bornean orangutan]]. It is an important refuge for many endemic forest species, including the [[Borneo elephant]], the [[eastern Sumatran rhinoceros]], the [[Bornean clouded leopard]], the [[Staurois tuberilinguis|Bornean rock frog]], the [[hose's palm civet]] and the [[dayak fruit bat]].<ref>{{cite web|url=http://wwf.panda.org/index.cfm?uNewsID=89620|title=Scientists find dozens of new species in Borneo rainforests|publisher=[[World Wide Fund for Nature]]|date=19 December 2006|access-date=26 May 2017}}</ref><ref>{{cite web|url=http://wwf.panda.org/what_we_do/where_we_work/borneo_forests/about_borneo_forests/borneo_animals/|title=Borneo wildlife|publisher=World Wide Fund for Nature|access-date=26 May 2017}}</ref> [[File:Borneo 19 May 2002.jpg|thumb|[[NASA]] satellite image of Borneo on 19 May 2002]] [[Borneo peat swamp forests|Peat swamp forests]] occupy the entire coastline of Borneo.<ref>{{cite book|author1=Kathy MacKinnon|author2=Gusti Hatta|title=Ecology of Kalimantan: Indonesian Borneo|url=https://books.google.com/books?id=FFXRAgAAQBAJ&pg=PT148|date=5 February 2013|publisher=Tuttle Publishing|isbn=978-1-4629-0505-8|pages=148β}}</ref> The soil of the peat swamp is comparatively infertile, while it is known to be the home of various bird species such as the [[hook-billed bulbul]], [[helmeted hornbill]] and [[rhinoceros hornbill]].<ref name=birds /> There are about 15,000 species of [[flowering plant]]s with 3,000 species of trees (267 species are [[dipterocarp]]s), 221 species of terrestrial mammals and 420 species of resident birds in Borneo.<ref>{{cite book|last=MacKinnon, K|title=The Ecology of Kalimantan|year=1998|publisher=[[Oxford University Press]]|location=London|display-authors=etal}}</ref> There are about 440 freshwater fish species in Borneo (about the same as Sumatra and Java combined).<ref>Nguyen, T.T.T., and S. S. De Silva (2006). "Freshwater Finfish Biodiversity and Conservation: An Asian Perspective", ''Biodiversity & Conservation'' 15(11): 3543β3568</ref> The [[Glyphis fowlerae|Borneo river shark]] is known only from the [[Kinabatangan River]].<ref>{{cite journal |last1=Li |first1=C. |last2=Corrigan |first2=S. |last3=Yang |first3=L. |last4=Straube |first4=N. |last5=Harris |first5=M. |last6=Hofreiter |first6=M. |last7=White |first7=W.T. |last8=Naylor |first8=G.J.P. |title=DNA capture reveals transoceanic gene flow in endangered river sharks |journal=Proceedings of the National Academy of Sciences |year=2015 |volume=112 |issue=43 |pages=13302β13307|doi=10.1073/pnas.1508735112 |pmid=26460025 |pmc=4629339|bibcode=2015PNAS..11213302L |doi-access=free }}</ref> In 2010, the [[World Wide Fund for Nature]] (WWF) stated that 123 species have been discovered in Borneo since the "[[Heart of Borneo]]" agreement was signed in 2007.<ref>{{cite web|url=https://www.wwf.org.uk/updates/scientists-discover-new-species-heart-borneo|title=Scientists discover new species in Heart of Borneo|publisher=World Wide Fund for Nature|date=22 April 2010|access-date=26 May 2017|archive-url=https://web.archive.org/web/20200228162702/https://www.wwf.org.uk/updates/scientists-discover-new-species-heart-borneo|archive-date=28 February 2020|url-status=dead}}</ref> [[File:Logging road East Kalimantan 2005.jpg|thumb|right|Logging road in [[East Kalimantan]], Indonesia]] The WWF has classified the island into seven distinct [[ecoregion]]s. Most are lowland regions:<ref name="Borneo forests">{{cite web|url=http://wwf.panda.org/about_our_earth/ecoregions/borneo_lowland_forests.cfm|title=Borneo Lowland & Montane Forests β A Global Ecoregion|publisher=World Wide Fund for Nature|access-date=26 May 2017|url-status=dead|archive-url=https://web.archive.org/web/20170715021114/http://wwf.panda.org/about_our_earth/ecoregions/borneo_lowland_forests.cfm|archive-date=15 July 2017}}</ref><ref>{{cite web|url=https://www.worldwildlife.org/ecoregions/im0104|title=Southeastern Asia: Indonesia, Malaysia, and Brunei|author=Colby Loucks|publisher=World Wide Fund for Nature|access-date=26 May 2017}}</ref><ref>{{cite web|url=https://www.worldwildlife.org/ecoregions/im0161|title=Sundaland heath forests|publisher=World Wide Fund for Nature|access-date=26 May 2017}}</ref> * [[Borneo lowland rain forests]] cover most of the island, with an area of {{convert|427500|km2}}. * [[Borneo peat swamp forests]] * ''[[Kerangas]]'' or Sundaland heath forests * [[Southwest Borneo freshwater swamp forests]] are found in the island's western and southern lowlands * [[Sunda Shelf mangroves]] * The [[Borneo montane rain forests]] lie in the central highlands of the island, above the {{convert|1000|m}} elevation. *The highest elevations of [[Mount Kinabalu]] are home to the [[Kinabalu montane alpine meadows]], a subalpine and alpine shrubland notable for its numerous endemic species, including many orchids. According to analysis of data from [[Global Forest Watch]],<ref>{{cite web|url=https://www.globalforestwatch.org|title=Global Forest Watch|publisher=World Resources Institute|access-date=14 August 2020}}</ref> the Indonesian portion of Borneo lost {{convert|10.7|e6ha|abbr=off}} of tree cover between 2002 and 2019, of which {{convert|4|e6ha|abbr=off}} was [[primary forest]], compared with Malaysian Borneo's {{convert|4.4|e6ha|abbr=off}} of tree cover loss and {{convert|1.9|e6ha|abbr=off}} of primary forest cover loss. As of 2020, Indonesian Borneo accounts for 72% of the island's tree cover, Malaysian Borneo 27%, and Brunei 1%. Primary forest in Indonesia accounts for 44% of Borneo's overall tree cover.<ref>{{cite web|url=https://rainforests.mongabay.com/borneo/|title=Borneo|publisher=Mongabay.com|access-date=14 August 2020}}</ref> ====Conservation issues==== {{see also|Deforestation in Borneo|1997 Indonesian forest fires|1997 Southeast Asian haze|2006 Southeast Asian haze|2013 Southeast Asian haze|2015 Southeast Asian haze|2016 Southeast Asian haze|2019 Southeast Asian haze}} [[File:Logging in Borneo (3540116932).jpg|thumb|Logging near [[Crocker Range National Park]]. Borneo has lost more than half of its rainforests in the past half century.<ref>{{cite news |title=What Causes Deforestation in Borneo and How Do We Stop It? |url=https://earth.org/deforestation-in-borneo/ |work=Earth.org |date=2 March 2023}}</ref>]] The island historically had extensive rainforest cover, but the area was reduced due to heavy [[logging]] by the [[Indonesia|Indonesian]] and [[Malaysia|Malaysian]] [[wood industry]], especially with the large demands of raw materials from [[developed country|industrial countries]] along with the conversion of forest lands for large-scale agricultural purposes.<ref name="Borneo forests"/> Half of the annual global [[tropical timber]] acquisition comes from Borneo. [[Palm oil]] plantations have been widely developed and are rapidly encroaching on the last remnants of primary rainforest.<ref>{{cite web|url=http://wwf.panda.org/what_we_do/where_we_work/borneo_forests/borneo_deforestation/|title=Threats to Borneo forests|publisher=World Wide Fund for Nature|access-date=26 May 2017|url-status=dead|archive-url=https://web.archive.org/web/20160525051215/http://wwf.panda.org/what_we_do/where_we_work/borneo_forests/borneo_deforestation/|archive-date=25 May 2016}}</ref> [[Wildfire|Forest fires]] since [[1997]], started by the locals to clear the forests for plantations were exacerbated by an exceptionally dry [[El NiΓ±o]] season, worsening the annual shrinkage of the rainforest.<ref>{{cite news|url=http://www.latimes.com/world/asia/la-fg-great-haze-explainer-20151021-htmlstory.html|title=Fires in Southeast Asia may be emitting more greenhouse gases than the entire U.S.|author=Jonathan Kaiman|newspaper=[[Los Angeles Times]]|date=21 October 2015|access-date=27 May 2017}}</ref> During these fires, hotspots were visible on [[Satellite imagery|satellite images]] and the resulting [[haze]] frequently affected [[Brunei]], [[Indonesia]] and [[Malaysia]]. The haze could also reach southern [[Thailand]], [[Cambodia]], [[Vietnam]] and the [[Philippines]] as evidenced on the [[2015 Southeast Asian haze]].<ref>{{cite web|url=http://www.eria.org/ERIA-DP-2015-82.pdf |archive-url=https://ghostarchive.org/archive/20221009/http://www.eria.org/ERIA-DP-2015-82.pdf |archive-date=9 October 2022 |url-status=live|title=Transboundary Haze Pollution Problem in Southeast Asia: Reframing ASEAN's Response|author=Apichal Sunchindah|publisher=Economic Research Institute for ASEAN and East Asia|date=December 2015|access-date=27 May 2017}}</ref> A study in [[2018]] found that [[Bornean orangutan]]s declined by 148,500 individuals from [[1999]] to [[2015]].<ref>{{cite news |last1=Solly |first1=Meilan |title=Borneo's Orangutan Population Plummeted by Half in 16 Years |url=https://www.smithsonianmag.com/smart-news/borneos-orangutan-population-plummeted-half-16-years-180968189/ |work=[[Smithsonian (magazine)|Smithsonian]] |date=16 February 2018}}</ref>
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