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==Systematics== {{cladogram|style=width:250px;font-size:85%;line-height:85%|caption=Subfamily relationships of Vespertilionidae<ref name="Hoofer 2003">{{cite journal| last1=Hoofer| first1= S. R.| last2= Bussche| first2= R. A. V. D.| date= 2003| title= Molecular phylogenetics of the chiropteran family Vespertilionidae| journal= Acta Chiropterologica| volume= 5| issue=1| pages= 1β63| doi= 10.3161/001.005.s101| doi-access= free| hdl= 11244/44678| hdl-access= free}}</ref> |1={{clade |label1='''Vespertilionidae''' |1={{clade |1=[[Vespertilioninae]] |2={{clade |1={{clade |1=[[Myotinae]] |2={{clade |1=[[Kerivoulinae]] |2=[[Murininae]] }} }} }} }} }} }} The four [[subfamilies]] of Vespertilionidae separate the presumably related taxa, tribes, and genera of extant and extinct taxa. The subfamilial treatments, based on morphological, geographical, and ecological comparisons have been recombined since the inclusion of the phylogenetic implications of [[molecular genetics]]; only the [[Murininae]] and [[Kerivoulinae]] have not been changed in light of genetic analysis.<ref name="Hoofer 2003"/> Subfamilies that were once recognized as valid, such as the [[Nyctophilinae]], are considered dubious, as molecular evidence suggests they are [[paraphyletic]] in their arrangements.<ref name="Hoofer 2003"/> Within the concept [[Yangochiroptera]], an acknowledged cladistic treatment, the closest relatives to the family are the free-tailed bats of family [[Molossidae]].<ref name="Hoofer 2003"/> The monotypic genus ''Tomopeas'', represented by the blunt-eared bat (''[[Tomopeas ravum]]''), is acknowledged as the potentially closest link between the Vespertilionidae and Molossidae, as it is the most [[basal (phylogenetics)|basal]] member of the Molossidae and has intermediate characteristics of both families.<ref name="Sudman 1994">{{cite journal| doi=10.2307/1382555| jstor=1382555| title=Familial Affinity of Tomopeas ravus (Chiroptera) Based on Protein Electrophoretic and Cytochrome b Sequence Data| journal=Journal of Mammalogy| volume=75| issue=2| pages=365| year=1994| last1=Sudman| first1=P. D| last2=Barkley| first2=L. J| last3=Hafner| first3=M. S}}</ref>
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