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===Morphology=== Tsetse flies can be seen as independent individuals in three forms: as third-[[instar]] larvae, pupae, and adults. Tsetse first becomes separate from their mothers during the third larval instar, during which they have the typical appearance of [[Fly#Maggots|maggots]]. However, this life stage is short, lasting at most a few hours, and is almost never observed outside of the laboratory. Tsetse next develops a hard external case, the puparium, and become pupae - small, hard-shelled oblongs with two distinctively small, dark lobes at the tail (breathing) end. Tsetse pupae are under {{ convert | 1 | cm | in | frac=4 }} long.<ref name="Jordan">{{Cite book |author=A. M. Jordan |year=1986 |title=Trypanosomaisis control and African rural development |publisher=London and New York: [[Longman]]}}</ref> Within the puparial shell, tsetse complete the last two larval instars and the pupal stage. At the end of the pupal stage, tsetse emerges as adult flies. The adults are relatively large flies, with lengths of {{ convert | 0.5 | - | 1.5 | cm | in | frac=8 }},<ref name="Jordan"/> and have a recognizable shape, or [[bauplan]], which makes them easy to distinguish from other flies. Tsetse have large heads, distinctly separated eyes, and unusual antennae. The [[Thorax (insect anatomy)|thorax]] is quite large, while the abdomen is wider, rather than elongated, and shorter than the wings. Four characteristics collectively separate adult tsetse from other kinds of flies: {|class="wikitable" |- | '''Proboscis''' || Tsetse have a distinct proboscis, a long and thin structure attached to the bottom of the head, pointing forward. || [[Image:tsetse head-proboscis.jpeg|center|70px|A photograph of the head of a tsetse illustrating the forward pointing proboscis]] |- | '''Folded wings''' || When at rest, tsetse fold their wings completely, one-on-top of the other. || [[Image:tsetse foldedWings.jpeg|center|200px|A photograph of the whole body of a tsetse illustrating the folded wings when at rest]] |- | '''Hatchet cell''' || The discal medial ("middle") cell of the wing has a characteristic hatchet shape, resembling a meat cleaver or a hatchet. || [[Image:tsetse hatchetCell.jpeg|center|140px|A photograph of the wing of a tsetse illustrating the hatchet shaped central cell]] |- | '''Branched arista hairs''' || The antennae have arista with hairs which are, themselves, branched. || [[Image:tsetse aristaHairs labeled.jpeg|center|140px|A photograph and diagram of the head of a tsetse illustrating the branched hairs of the antenna's arista]] |}
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