Editing Pteranodon (section)

===A toothless pterosaur===
[[File:Pteranodon longiceps YPM1177.jpg|thumb|left|Short-crested ''P. longiceps'' holotype specimen YPM1177, now interpreted as a female individual]]

While the first ''Pteranodon'' wing bones were collected by Marsh and Cope in the early 1870s, the first ''Pteranodon'' skull was found on May 2, 1876, along the [[Smoky Hill River]] in [[Wallace County, Kansas|Wallace County]] (now Logan County), Kansas, USA, by [[Samuel Wendell Williston]], a fossil collector working for Marsh.<ref name= "bennett2000">Bennett, S.C. (2000). "Inferring stratigraphic position of fossil vertebrates from the Niobrara Chalk of western Kansas." ''Current Research in Earth Sciences: Kansas Geological Survey Bulletin'', '''244'''(Part 1): 26 pp.</ref> A second, smaller skull soon was discovered as well. These skulls showed that the North American pterosaurs were different from any European species, in that they lacked teeth and had bony crests on their skulls. Marsh recognized this major difference, describing the specimens as "distinguished from all previously known genera of the order Pterosauria by the entire absence of teeth." Marsh recognized that this characteristic warranted a new genus, and he coined the name ''Pteranodon'' ("wing without tooth") in 1876. Marsh reclassified all the previously named North American species from ''Pterodactylus'' to ''Pteranodon''. He considered the smaller skull to belong to ''Pteranodon occidentalis'', based on its size. Marsh classified the larger skull, YPM 1117, in the new species ''Pteranodon longiceps'', which he thought to be a medium-sized species in between the small ''P. occidentalis'' and the large ''P. ingens''.<ref name=marsh1876a>{{Cite journal | last = Marsh| first= O.C. | year = 1876a | title = Notice of a new sub-order of Pterosauria | url = https://zenodo.org/record/1450032| journal = American Journal of Science |series=Series 3 | volume = 11 | issue = 65| pages = 507–509 | doi=10.2475/ajs.s3-11.66.507| bibcode= 1876AmJS...11..507M | s2cid = 130203580 }}</ref><ref name=bennett1994/> Marsh also named several additional species: '''''Pteranodon comptus''''' and ''[[Pteranodon nanus]]'' were named for fragmentary skeletons of small individuals, while ''Pteranodon gracilis'' was based on a wing bone that he mistook for a pelvic bone. He soon realized his mistake, and re-classified that specimen again into a separate genus, which he named ''[[Nyctosaurus]]''. ''P. nanus'' was also later recognized as a ''Nyctosaurus'' specimen.<ref name=marsh1976b>{{Cite journal |  last= Marsh | first= O.C. | year = 1876b | title = Principal characters of American pterodactyls | url = https://zenodo.org/record/1450034| journal = American Journal of Science |series=Series 3 | volume = 12 | issue = 72| pages = 479–480 | doi=10.2475/ajs.s3-12.72.479| bibcode= 1876AmJS...12..479M | s2cid = 131057784 }}</ref><ref name=bennett1994/>

In 1892, Samuel Williston examined the question of ''Pteranodon'' classification. He noticed that, in 1871, Seeley had mentioned the existence of a partial set of toothless pterosaur jaws from the [[Cambridge Greensand]] of [[England]], which he named ''[[Ornithostoma]]''. Because the primary characteristic Marsh had used to separate ''Pteranodon'' from other pterosaurs was its lack of teeth, Williston concluded that "Ornithostoma" must be considered the senior synonym of ''Pteranodon''. However, in 1901, Pleininger pointed out that "Ornithostoma" had never been scientifically described or even assigned a species name until Williston's work, and therefore had been a ''[[nomen nudum]]'' and could not beat out ''Pteranodon'' for naming priority. Williston accepted this conclusion and went back to calling the genus ''Pteranodon''.<ref name= bennett1994/> However, both Williston and Pleininger were incorrect, because unnoticed by both of them was the fact that, in 1891, Seeley himself had finally described and properly named ''Ornithostoma'', assigning it to the species ''O. sedgwicki''. In the 2010s, more research on the identity of ''Ornithostoma'' showed that it was probably not ''Pteranodon'' or even a close relative, but may in fact have been an [[Azhdarchoidea|azhdarchoid]], a different type of toothless pterosaur.<ref name=Averianov2012>{{cite journal | last1 = Averianov | first1 = A.O. | year = 2012 | title = ''Ornithostoma sedgwicki'' – valid taxon of azhdarchoid pterosaurs | journal = Proceedings of the Zoological Institute RAS | volume = 316 | issue = 1 | pages = 40–49| doi = 10.31610/trudyzin/2012.316.1.40 | s2cid = 67809186 | doi-access = free }}</ref>