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=== Genetic studies === A 2015 study by Larmuseau et al. investigated the paternal genetic landscape of populations along the Bight of Benin, focusing on four groups from present-day Benin: the Fon, Yoruba, Bariba, and Dendi. Using Y-chromosomal markers, the research revealed significant genetic diversity among these populations, reflecting complex historical migrations and interactions. The study found that the Beninese Yoruba exhibit notable genetic differentiation not only from neighboring groups but also from the Yoruba in Nigeria, suggesting distinct paternal lineages and historical developments. This diversity underscores the intricate demographic history of the region, shaped by various migration events and cultural exchanges.<ref>{{cite journal |last1=Larmuseau |first1=Maarten H. D. |last2=Vessi |first2=Andrea |last3=Jobling |first3=Mark A. |last4=Van Geystelen |first4=Anneleen |last5=Primativo |first5=Giuseppina |last6=Biondi |first6=Gianfranco |last7=Martínez-Labarga |first7=Cristina |last8=Ottoni |first8=Claudio |last9=Decorte |first9=Ronny |last10=Rickards |first10=Olga |title=The Paternal Landscape along the Bight of Benin – Testing Regional Representativeness of West-African Population Samples Using Y-Chromosomal Markers |journal=PLOS ONE |volume=10 |issue=11 |pages=e0141510 |year=2015 |doi=10.1371/journal.pone.0141510 |doi-access=free |pmid=26544036 |pmc=4636292 |bibcode=2015PLoSO..1041510L }}</ref>
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