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=== Taxonomy === About 88% of [[amphibian]] species are [[Biological classification|classified]] in the [[Order (biology)|order]] Anura.<ref name="Pou92">{{cite book |last1=Pough |first1=F. H. |last2=Andrews |first2=R. M. |last3=Cadle |first3=J. E. |last4=Crump |first4=M. L. |last5=Savitsky |first5=A. H. |last6=Wells |first6=K. D. |title=Herpetology |edition=third |year=2003 |publisher=Benjamin Cummings |isbn=978-0-13-100849-6 }}</ref> These include over 7,700 species<ref name="AOTW"/> in 59 [[Family (biology)|families]], of which the [[Hylidae]] (1062 spp.), [[Strabomantidae]] (807 spp.), [[Microhylidae]] (758 spp.), and [[True toad|Bufonidae]] (657 spp.) are the [[Species richness|richest in species]].<ref name=amdb>{{cite web |url=http://research.amnh.org/vz/herpetology/amphibia/Amphibia/Anura |title=Anura Fischer von Waldheim, 1813 |last=Frost |first=Darrel R. |year=2023 |work=Amphibian Species of the World: an Online Reference. Version 6.2 |publisher=American Museum of Natural History |access-date=November 12, 2024}}</ref> [[File:Bombina bombina 1 (Marek Szczepanek) tight crop.jpg|thumb|left|alt=Dark-coloured toad facing left|[[European fire-bellied toad]] (''Bombina bombina'')]] The Anura include all modern frogs and any [[fossil]] species that fit within the anuran definition. The characteristics of anuran adults include: 9 or fewer presacral vertebrae, the presence of a urostyle formed of fused vertebrae, no tail, a long and forward-sloping ilium, shorter fore limbs than hind limbs, [[Radius (bone)|radius]] and [[ulna]] fused, [[tibia]] and [[fibula]] fused, elongated [[ankle bone]]s, absence of a prefrontal bone, presence of a [[hyoid bone|hyoid plate]], a [[Mandible|lower jaw]] without teeth (with the exception of ''[[Gastrotheca guentheri]]'') consisting of three pairs of bones (angulosplenial, dentary, and mentomeckelian, with the last pair being absent in [[Pipoidea]]),<ref>Duellman, William E. {{Google books|CzxVvKmrtIgC|Biology of Amphibians|page=319}}</ref> an unsupported tongue, [[lymph]] spaces underneath the skin, and a muscle, the protractor lentis, attached to the [[Lens (anatomy)|lens of the eye]].<ref name=ToLAnura>{{cite web |url=http://www.tolweb.org/Anura/16963 |title=Anura |author=Cannatella, David |author-link=David C. Cannatella |date=January 11, 2008 |publisher=Tree of Life web project |access-date=August 8, 2012}}</ref> The anuran larva or tadpole has a single central respiratory [[Spiracle (vertebrates)|spiracle]] and mouthparts consisting of [[keratin]]ous beaks and [[Denticle (tooth feature)|denticles]].<ref name=ToLAnura/> [[File:Atelopus zeteki1.jpg|thumb|alt=Panamanian golden frog|[[Panamanian golden frog]] (''Atelopus zeteki'')]] Frogs and toads are broadly classified into three suborders: [[Archaeobatrachia]], which includes four families of primitive frogs; [[Mesobatrachia]], which includes five families of more evolutionary intermediate frogs; and [[Neobatrachia]], by far the largest group, which contains the remaining families of modern frogs, including most common species throughout the world. The suborder Neobatrachia is further divided into the two superfamilies [[Hyloidea]] and [[Ranoidea]].<ref>{{cite journal |last=Ford |first=L.S. |author2=Cannatella, D. C. |year=1993 |title=The major clades of frogs |journal=Herpetological Monographs |volume=7 |pages=94โ117 |doi=10.2307/1466954 |jstor=1466954}}</ref> This classification is based on such [[Morphology (biology)|morphological]] features as the number of vertebrae, the structure of the [[pectoral girdle]], and the morphology of tadpoles. While this classification is largely accepted, relationships among families of frogs are still debated.<ref>{{cite journal |last=Faivovich |first=J. |author2=Haddad, C. F. B. |author3=Garcia, P. C. A. |author4=Frost, D. R. |author5=Campbell, J. A. |author6=Wheeler, W. C. |title=Systematic review of the frog family Hylidae, with special reference to Hylinae: Phylogenetic analysis and revision |journal=Bulletin of the American Museum of Natural History |volume=294 |pages=1โ240 |doi=10.1206/0003-0090(2005)294[0001:SROTFF]2.0.CO;2 |year=2005 |citeseerx=10.1.1.470.2967 |s2cid=83925199}}</ref> Some species of anurans [[hybrid (biology)|hybridise]] readily. For instance, the [[edible frog]] (''Pelophylax esculentus'') is a hybrid between the [[pool frog]] (''P. lessonae'') and the [[marsh frog]] (''P. ridibundus'').<ref>{{cite web |url=http://amphibiaweb.org/cgi-bin/amphib_query?where-genus=Pelophylax&where-species=esculentus |title=''Pelophylax esculentus'' |author=Kuzmin, S. L. |date=November 10, 1999 |access-date=October 12, 2012}}</ref> The fire-bellied toads ''Bombina bombina'' and ''[[Yellow-bellied toad|B. variegata]]'' are similar in forming hybrids. These are less fertile than their parents, giving rise to a [[hybrid zone]] where the hybrids are prevalent.<ref>{{cite web |url=http://edoc.ub.uni-muenchen.de/1521/1/Koehler_Sonja.pdf |archive-url=https://ghostarchive.org/archive/20221009/http://edoc.ub.uni-muenchen.de/1521/1/Koehler_Sonja.pdf |archive-date=October 9, 2022 |url-status=live |title=Mechanisms for partial reproductive isolation in a ''Bombina'' hybrid zone in Romania |last=Kรถhler |first=Sonja |year=2003 |work=Dissertation for thesis |access-date=June 5, 2012}}</ref> {{clear left}}
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