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==Classification== Traditionally, five subfamilies had been distinguished within the Felidae based on [[Phenotype|phenotypical]] features: the [[Pantherinae]], the [[Felinae]], the Acinonychinae,<ref name=Pocock_1917 /> and the extinct [[Machairodontinae]] and [[Proailurinae]].<ref name="McKenna & Bell">{{cite book |last1=McKenna |first1=M. C. |last2=Bell |first2=S. K. |title=Classification of Mammals |date=2000 |publisher=Columbia University Press |isbn=9780231110136 |page=230 |chapter=Family Felidae Fischer de Waldheim, 1817:372. Cats |chapter-url=https://books.google.com/books?id=zS7FZkzIw-cC&pg=PA230 |access-date=2020-12-31 |archive-date=2021-04-19 |archive-url=https://web.archive.org/web/20210419161520/https://books.google.com/books?id=zS7FZkzIw-cC&pg=PA230 |url-status=live}}</ref> Acinonychinae used to only contain the genus ''[[Acinonyx]]'' but this genus is now within the Felinae subfamily.<ref name=CatSG2017 /> ===Phylogeny=== The following cladogram based on Piras et al. (2013) depicts the phylogeny of basal living and extinct groups.<ref name="PirasMaiorino2013">{{cite journal |last1=Piras |first1=P. |last2=Maiorino |first2=L. |last3=Teresi |first3=L. |last4=Meloro |first4=C. |last5=Lucci |first5=F. |last6=Kotsakis |first6=T. |last7=Raia |first7=P. |title=Bite of the Cats: Relationships between Functional Integration and Mechanical Performance as Revealed by Mandible Geometry |journal=Systematic Biology |volume=62 |issue=6 |date=2013 |pages=878β900 |issn=1063-5157 |doi=10.1093/sysbio/syt053 |pmid=23925509|doi-access=free}}</ref> {{clade |style=font-size:85%;line-height:85%; |label1='''Felidae''' |1={{clade |style1=background-color:#eeccFF; |label1=β ''[[Proailurus]]'' |1={{clade |caption=Proailurinae |captionstyle=text-align:middle;font-weight:bold;color:purple;padding:2px 10px 1px 1px; |1=β ''Proailurus bourbonnensis'' |2=β ''Proailurus lemanensis'' |3=β ''Proailurus major'' }} |label2=''"Pseudaelurus"''|sublabel2=(grade) |2={{clade |style1=background-color:#B0CEFF; |1={{clade |caption=Pseudaelurus lineage |captionstyle=text-align:right;font-weight:bold;color:blue;padding:2px 10px 1px 1px; |label1=β ''[[Pseudaelurus]]'' |1={{clade |1=β ''Pseudaelurus quadridentatus'' |2=β ''Pseudaelurus cuspidatus'' |3=β ''Pseudaelurus guangheesis'' }} |2=β '''[[Machairodontinae]]''' [[File:Smilodon_fatalis.jpg|50px]] }} |label2=β ''[[Hyperailurictis]]''|style2=background-color:#FFB6B6; |2={{clade |caption=Hyperailurictis lineage |captionstyle=text-align:right;font-weight:bold;color:#882277;padding:2px 10px 1px 1px; |1=β ''Hyperailurictis intrepidus'' |2=β ''Hyperailurictis marshi'' |3=β ''Hyperailurictis stouti'' |4=β ''Hyperailurictis validus'' |5=β ''Hyperailurictis skinneri'' }} |label3=β ''[[Sivaelurus]]''|style3=background-color:#ddddFF; |3=β ''Sivaelurus chinjiensis'' |style4=background-color:#CCFF99; |4={{clade |caption=Styriofelis lineage |captionstyle=text-align:right;font-weight:bold;color:green;padding:2px 10px 1px 1px; |label1=β ''[[Styriofelis]]'' |1={{clade |1=β ''Styriofelis turnauensis'' |2=β ''Styriofelis romieviensis'' }} |label2='''Felinae'''|sublabel2=''sensu lato'' |2={{clade |1='''[[Felinae]]''' [[File:Anatomie_descriptive_et_comparative_du_chat_(1845)_Pl-I_(white_background_%26_colourised).jpg|50px]] |2={{clade |label1=β ''[[Miopanthera]]'' |1={{clade |1=β ''Miopanthera lorteti'' |2=β ''Miopanthera pamiri'' }} |2='''[[Pantherinae]]''' [[File:Stamp-russia2014-save-russian-cats-(snow_leopard).png|50px]] }} }} }} }} }} }} The phylogenetic relationships of living felids are shown in the following cladogram:<ref name="Li_al2016">{{cite journal |last1=Li |first1=G. |last2=Davis |first2=B. W. |last3=Eizirik |first3=E. |last4=Murphy |first4=W. J. |date=2016 |title=Phylogenomic evidence for ancient hybridization in the genomes of living cats (Felidae) |journal=Genome Research |volume=26 |issue=1 |pages=1β11 |doi=10.1101/gr.186668.114 |pmc=4691742 |pmid=26518481}}</ref> {{clade |style=font-size:85%;line-height:85%;background-color:#eeeeFF;border:1px solid darkred; |thickness=2 |label1='''Felidae''' |1={{clade |thickness=2 |caption=Panthera lineage |captionstyle=background-color:#eeccFF;color:#880088;text-align:right;font-weight:bold;padding:2px 10px 1px 1px; |color1=red |style1=background-color:#eeccFF; |label1='''[[Pantherinae]]''' |1={{clade |thickness=2 |color=red |label1=''[[Neofelis]]'' |1={{clade |thickness=2 |color=red |1=[[Sunda clouded leopard]] (''N. diardi'') |2=[[Clouded leopard]] (''N. nebulosa'') }} |label2=''[[Panthera]]'' |2={{clade |thickness=2 |color=red |1={{clade |thickness=2 |color=red |1=[[Tiger]] (''P. tigris'') |2=[[Snow leopard]] (''P. uncia'') }} |2={{clade |thickness=2 |color=red |1=[[Jaguar]] (''P. onca'') |2={{clade |thickness=2 |color=red |1=[[Lion]] (''P. leo'') |2=[[Leopard]] (''P. pardus'') }} }} }} }} |label2='''[[Felinae]]''' |2={{clade |thickness=2 |caption=Caracal lineage |captionstyle=background-color:#eeFFFF;color:darkcyan;text-align:right;font-weight:bold;padding:1px 10px 1px 1px; |color1=darkcyan |style1=background-color:#eeFFFF; |1={{clade |thickness=2 |color=darkcyan |label1=''[[Leptailurus]]'' |1=[[Serval]] (''L. serval'') |label2=''[[Caracal (genus)|Caracal]]'' |2={{clade |thickness=2 |color=darkcyan |1=[[African golden cat]] (''C. aurata'') |2=[[Caracal]] (''C. caracal'') }} }} |2={{clade |thickness=2 |caption=Ocelot lineage |captionstyle=background-color:#ccFFcc;color:darkgreen;text-align:right;font-weight:bold;padding:1px 10px 1px 1px; |color1=green |style1=background-color:#ccFFcc; |label1=''[[Leopardus]]'' |1={{clade |thickness=2 |color=green |1={{clade |thickness=2 |color=green |1=[[Andean mountain cat]] (''L. jacobita'') |2={{clade |thickness=2 |color=green |1=[[Ocelot]] (''L. pardalis'') |2=[[Margay]] (''L. wiedii'') }} }} |2={{clade |thickness=2 |color=green |1={{clade |thickness=2 |color=green |1=[[Pampas cat]] (''L. colocola'') }} |2={{clade |thickness=2 |color=green |1=[[Oncilla]] (Northern tiger cat, ''L. tigrina'') |2={{clade |thickness=2 |color=green |1=[[Southern tiger cat]] (''L. guttulus'') |2={{clade |thickness=2 |color=green |1=[[Geoffroy's cat]] (''L. geoffroyi'') |2=[[Kodkod]] (''L. guigna'') }} }} }} }} }} |2={{clade |thickness=2 |1={{clade |thickness=2 |caption=Bay cat lineage |captionstyle=background-color:#ddddFF;color:blue;text-align:right;font-weight:bold;padding:1px 10px 2px 1px; |color1=blue |style1=background-color:#ddddFF; |1={{clade |thickness=2 |color=blue |label1=''[[Pardofelis]]'' |1=[[Marbled cat]] (''P. marmorata'') |label2=''[[Catopuma]]'' |2={{clade |thickness=2 |color=blue |1=[[Bay cat]] (''C. badia'') |2=[[Asian golden cat]] (''C. temminckii'') }} }} |2={{clade |thickness=2 |captionx=Lynx lineage |captionstyle=background-color:#FFFFaa;color:darkorange;text-align:right;font-weight:bold;padding:1px 10px 1px 1px; |color1=darkorange |style1=background-color:#ffffaa; |label1=''[[Lynx]]'' |1={{clade |thickness=2 |color=darkorange |caption=Lynx lineage |captionstyle=background-color:#FFFFaa;color:darkorange;text-align:right;font-weight:bold;padding:1px 10px 1px 1px; |1=[[Bobcat]] (''L. rufus'') |2={{clade |thickness=2 |color=darkorange |1=[[Canada lynx]] (''L. canadensis'') |2={{clade |thickness=2 |color=darkorange |1=[[Eurasian lynx]] (''L. lynx'') |2=[[Iberian lynx]] (''L. pardinus'') }} }} }} }} }} |2={{clade |thickness=2 |caption=Puma lineage |captionstyle=background-color:#ffdd66;color:#dd4400;text-align:right;font-weight:bold;padding:1px 10px 1px 1px; |color1=#dd4400 |style1=background-color:#ffdd66; |1={{clade |thickness=2 |color=#dd4400 |label1=''[[Acinonyx]]'' |1=[[Cheetah]] (''A. jubatus'') |2={{clade |thickness=2 |color=#dd4400 |label1=''[[Herpailurus]]'' |1=[[Jaguarundi]] (''H. yagouaroundi'') |label2=''[[Puma (genus)|Puma]]'' |2=[[Cougar]] (''P. concolor'') }} }} |2={{clade |thickness=2 |caption=Leopard cat lineage |captionstyle=background-color:#ffdddd;color:#aa3322;text-align:right;font-weight:bold;padding:1px 10px 2px 1px; |color1=#aa3322 |style1=background-color:#FFdddd; |1={{clade |thickness=2 |color=#aa3322 |label1=''[[Otocolobus]]'' |1=[[Pallas's cat]] (''O. manul'') |label2=''[[Prionailurus]]'' |2={{clade |thickness=2 |color=#aa3322 |1=[[Rusty-spotted cat]] (''P. rubiginosus'') |2={{clade |thickness=2 |color=#aa3322 |1=[[Flat-headed cat]] (''P. planiceps'') |2={{clade |thickness=2 |color=#aa3322 |1=[[Fishing cat]] (''P. viverrinus'') |2={{clade |thickness=2 |color=#aa3322 |1=[[Leopard cat]] (''P. bengalensis'') |2=[[Sunda leopard cat]] (''P. javanensis'') }} }} }} }} }} |color2=#882277 |style2=background-color:#ffbbdd; |label2=''[[Felis]]'' |2={{clade |thickness=2 |color=#882277 |caption=Domestic cat lineage |captionstyle=background-color:#ffbbdd;color:#882277;text-align:right;font-weight:bold;padding:1px 10px 10px 1px; |1=[[Jungle cat]] (''F. chaus'') |2={{clade |thickness=2 |color=#882277 |1=[[Black-footed cat]] (''F. nigripes'') |2={{clade |thickness=2 |color=#882277 |1=[[Sand cat]] (''F. margarita'') |label2=[[wildcat]]s |2={{clade |thickness=2 |color=#882277 |1={{clade |thickness=2 |color=#882277 |1=[[Chinese mountain cat]] (''F. bieti'') |2=[[African wildcat]] (''F. lybica'') }} |2={{clade |thickness=2 |color=#882277 |1=[[Cat|Domestic cat]] (''F. catus'') |2=[[European wildcat]] (''F. silvestris'') }} }} }} }} }} }} }} }} }} }} }} }}
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