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== Extinctions by severity == {{Main|List of extinction events}} Extinction events can be tracked by several methods, including geological change, ecological impact, extinction vs. origination ([[speciation]]) rates, and most commonly diversity loss among [[Taxonomy|taxonomic]] units. Most early papers used [[Family (biology)|families]] as the unit of taxonomy, based on compendiums of marine animal families by [[Jack Sepkoski|Sepkoski]] (1982, 1992).<ref name="Sepkoski_1982" /><ref name="Sepkoski_1992" /> Later papers by Sepkoski and other authors switched to [[genera]], which are more precise than families and less prone to taxonomic bias or incomplete sampling relative to species.<ref name="Sepkoski_2002" /> These are several major papers estimating loss or ecological impact from fifteen commonly-discussed extinction events. Different methods used by these papers are described in the following section. The "Big Five" mass extinctions are bolded. {| class="wikitable sortable" style="text-align: center;" |+Extinction proportions (diversity loss) of marine genera or ecological impact in estimates of mass extinction severity !rowspan=2|Extinction name !rowspan=2|Age<br>([[Myr|Ma]]) !rowspan=2|Sepkoski (1996)<ref name="Sepkoski_1996" /><br>Multiple-interval<br>genera !rowspan=2|Bambach<br>(2006)<ref name="Bambach_2006" /> !colspan=2|McGhee ''et al''. (2013)<ref name="McGhee_2013" /> !rowspan=2|Stanley (2016)<ref name="Stanley_2016" /> |- !Taxonomic<br>loss !Ecological<br>ranking |- |'''[[Late Ordovician mass extinction|Late Ordovician]] ([[Ashgill (age)|Ashgillian]] / [[Hirnantian]])''' |445–444 |~49% |57%{{ref label|total|d|d}}<br>(40%, 31%){{ref label|2pulse|e|e}} |52% |7 |42–46% |- |[[Lau event]] ([[Ludfordian]]) |424 |~23% | – |9% |9 | – |- |[[Kačák Event]] ([[Eifelian]]) |388~ |~24%{{ref label|Sepgraph|a|a}} | – |32% |9 | – |- |[[Taghanic Event]] ([[Givetian]]) |384~ |~30%{{ref label|Sepgraph|a|a}} |28.5% |36% |8 | – |- |'''[[Late Devonian extinction|Late Devonian]]/[[Kellwasser event]] ([[Frasnian]])''' |372 |~35% |34.7% |40% |4 |16–20% |- |End-Devonian/[[Hangenberg event]] ([[Famennian]]) |359 |~28%{{ref label|Sepgraph|a|a}} |31% |50% |7 |<13%{{ref label|noS16|f|f}} |- |[[Serpukhovian#Serpukhovian extinction|Serpukhovian]] |330–325~ |~23% |31% |39% |6 |13–15% |- |[[Capitanian mass extinction event|Capitanian]] |260 |~47%{{ref label|Cap|b|b}} |48% |25% |5 |33–35% |- |'''[[Permian–Triassic extinction event|Permian–Triassic]] ([[Changhsingian]])''' |252 |~58% |55.7% |83% |1 |62% |- |'''[[Triassic–Jurassic extinction event|Triassic–Jurassic]] ([[Rhaetian]])''' |201 |~37%{{ref label|Norian|c|c}} |47%{{ref label|Norian|c|c}} |73% |3 |N/A{{ref label|timing|g|g}} |- |[[Toarcian Oceanic Anoxic Event|Pliensbachian-Toarcian]] |186–178 |~14% |25%, 20%{{ref label|2pulse|e|e}} | – | – | – |- |[[Tithonian#Tithonian extinction|End-Jurassic]] (Tithonian) |145 |~18% |20% | – | – | – |- |[[Cenomanian-Turonian boundary event|Cenomanian-Turonian]] |94 |~15% |25% | – | – | – |- |'''[[Cretaceous–Paleogene extinction event|Cretaceous–Paleogene]] ([[Maastrichtian]])''' |66 |~39% |40–47% |40% |2 |38–40% |- |[[Eocene–Oligocene extinction event|Eocene–Oligocene]] |34 |~11% |15.6% | – | – | – |} {{note label|Sepgraph|a|a}} Graphed but not discussed by Sepkoski (1996), considered continuous with the Late Devonian mass extinction<br>{{note label|Cap|b|b}} At the time considered continuous with the end-Permian mass extinction<br>{{note label|Norian|c|c}} Includes late [[Norian]] time slices<br>{{note label|total|d|d}} Diversity loss of both pulses calculated together<br>{{note label|2pulse|e|e}} Pulses extend over adjacent time slices, calculated separately<br>{{note label|noS16|f|f}} Considered ecologically significant, but not analyzed directly<br>{{note label|timing|g|g}} Excluded due to a lack of consensus on Late Triassic chronology
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