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== Differences between evergreen and deciduous species == Evergreen and deciduous species vary in a range of morphological and physiological characters. Generally, broad-leaved evergreen species have thicker leaves than deciduous species, with a larger volume of [[Ground tissue|parenchyma]] and air spaces per unit leaf area.<ref>{{Cite journal |last=Villar |first=Rafael |last2=Ruiz-Robleto |first2=Jeannete |last3=Ubera |first3=JosΓ© Luis |last4=Poorter |first4=Hendrik |date=October 2013 |title=Exploring variation in leaf mass per area (LMA) from leaf to cell: An anatomical analysis of 26 woody species |journal=American Journal of Botany |volume=100 |issue=10 |pages=1969β1980 |doi=10.3732/ajb.1200562 |pmid=24107583}}</ref> They have larger leaf biomass per unit leaf area, and hence a lower [[specific leaf area]]. Construction costs do not differ between the groups.{{citation needed|date=February 2023}} Evergreens have generally a larger fraction of total plant biomass present as leaves (LMF),<ref>{{Cite journal |last=Poorter |first=Hendrik |last2=Jagodzinski |first2=Andrzej M. |last3=Ruiz-Peinado |first3=Ricardo |last4=Kuyah |first4=Shem |last5=Luo |first5=Yunjian |last6=Oleksyn |first6=Jacek |last7=Usoltsev |first7=Vladimir A. |last8=Buckley |first8=Thomas N. |last9=Reich |first9=Peter B. |last10=Sack |first10=Lawren |date=2015 |title=How does biomass distribution change with size and differ among species? An analysis for 1200 plant species from five continents |journal=New Phytologist |volume=208 |issue=3 |pages=736β749 |bibcode=2015NewPh.208..736P |doi=10.1111/nph.13571 |pmc=5034769 |pmid=26197869}}</ref> but they often have a lower rate of photosynthesis.
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