Editing Dhole (section)

=== Admixture with the African wild dog ===
In 2018, [[whole genome sequencing]] was used to compare all members (apart from the black-backed and side-striped jackals) of the genus ''Canis'', along with the dhole and the [[African wild dog]] (''Lycaon pictus''). There was strong evidence of ancient [[genetic admixture]] between the dhole and the African wild dog. Today, their ranges are remote from each other; however, during the [[Pleistocene]] era the dhole could be found as far west as Europe. The study proposes that the dhole's distribution may have once included the [[Middle East]], from where it may have admixed with the African wild dog in [[North Africa]]. However, there is no evidence of the dhole having existed in the Middle East nor North Africa,<ref name=gopalakrishnan2018>{{cite journal |doi=10.1016/j.cub.2018.08.041 |pmid=30344120 |pmc=6224481 |title=Interspecific gene flow shaped the evolution of the Genus ''Canis'' |journal=Current Biology |volume=28 |issue=21 |pages=3441–3449.e5 |year=2018|last1=Gopalakrishnan|first1=S. |last2=Sinding|first2=M.-H.S. |last3=Ramos-Madrigal|first3=J. |last4= Niemann |first4=J. |last5= Samaniego Castruita|first5=J.A. |last6=Vieira|first6=F.G. |last7= Carøe|first7=C. |last8=Montero |first8= M.d.M. |last9=Kuderna| first9=L. |last10= Serres|first10=A. |last11= González-Basallote |first11=V.M. |last12=Liu|first12=Y.-H. |last13=Wang |first13=G.-D. |last14=Marques-Bonet |first14=T. |last15=Mirarab |first15=S. |last16=Fernandes|first16=C. |last17= Gaubert|first17=P. |last18= Koepfli|first18=K.-P. |last19=Budd |first19=J. |last20= Rueness| first20= E.K. |last21=Heide-Jørgensen| first21= M.P. |last22= Petersen |first22=B. |last23=Sicheritz-Ponten|first23=T. |last24= Bachmann| first24=L. |last25= Wiig|first25=Ø. |last26=Hansen |first26= A.J. |last27= Gilbert| first27= M.T.P. |bibcode=2018CBio...28E3441G | display-authors= 3}}</ref> though the ''Lycaon'' was present in Europe during the Early Pleistocene, with its last record in the region dating to 830,000 years ago. Genetic evidence from the Sardinan dhole suggests that both Sardinian and modern dholes (which are estimated to have split from each other around 900,000 years ago) share ancestry from the ''Lycaon'' lineage, but this ancestry is significantly higher in modern dholes than in the Sardinian dhole.<ref name=":0"/>
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