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==Distribution and diversity== Odonates are found on all the continents except Antarctica.<ref name=Nilsson>{{cite book|author=Nilsson, Anders |title=Aquatic insects of North Europe: A taxonomic handbook |url=https://books.google.com/books?id=mlAgAQAAMAAJ |year=1997 |publisher=Apollo Books |isbn=978-87-88757-07-1 |page=14}}</ref> Although some species of dragonfly have wide distributions, damselflies tend to have smaller ranges. Most odonates breed in fresh-water; a few damselflies in the family Caenagrionidae breed in brackish water (and a single dragonfly species breeds in seawater).<ref>{{cite journal|title=Observations and Experiments on Dragon-Flies in Brackish Water |first=Raymond C.|last= Osburn |journal=The American Naturalist|volume=40|issue=474|year= 1906|pages= 395β399|doi=10.1086/278632 |url=https://zenodo.org/record/1431341|doi-access=free|bibcode=1906ANat...40..395O }}</ref><ref>{{cite journal|title=Adaptations of Nymphs of a Marine Dragonfly, Erythrodiplax berenice, to Wide Variations in Salinity|first=William A. |last=Dunson|journal=Physiological Zoology|volume=53|issue=4|year=1980|pages=445β452|doi=10.1086/physzool.53.4.30157882 |s2cid=87698039 }}</ref> Dragonflies are more affected by pollution than are damselflies. The presence of odonates indicates that an ecosystem is of good quality. The most species-rich environments have a range of suitable microhabitats, providing suitable water bodies for breeding.<ref name=UCMP>{{cite web |title=Introduction to the Odonata |url=http://www.ucmp.berkeley.edu/arthropoda/uniramia/odonatoida.html |publisher=UCMP Berkeley |access-date=13 March 2015 |archive-url=https://web.archive.org/web/20170827225730/http://www.ucmp.berkeley.edu/arthropoda/uniramia/odonatoida.html |archive-date=27 August 2017 |url-status=dead }}</ref><ref name=diversity /> Although most damselflies live out their lives within a short distance of where they were hatched, some species, and some individuals within species, disperse more widely. [[Ischnura|Forktails]] in the family [[Coenagrionidae]] seem particularly prone to do this, large male [[boreal bluet]]s (''Enallagma boreale'') in British Columbia often migrating, while smaller ones do not.<ref name=Acorn/> These are known to leave their waterside habitats, flying upwards till lost from view, and presumably being dispersed to far off places by the stronger winds found at high altitudes.<ref name=Acorn/> In this way they may appear in a locality where no damselflies were to be seen the day before. [[Rambur's forktail]] (''Ischnura ramburii'') has been found, for example, on [[Oil platform|oil rigs]] far out in the [[Gulf of Mexico]].<ref name=Paulson2011/> The distribution and diversity of damselfly species in the [[biogeographical]] regions is summarized here. (There are no damselflies in the [[Antarctic]].) Note that some species are widespread and occur in multiple regions.<ref name=diversity /> {| class="wikitable sortable collapsible collapsed" |- style="font-size:86%" ! Family ! [[Indomalaya|Oriental]] ! [[Neotropical]] ! [[Australasia]]n ! [[Afrotropical]] ! [[Palaearctic]] ! [[Nearctic]] ! [[Oceania|Pacific]] ! World |- | [[Hemiphlebiidae]]||||||1||||||||||1 |- | [[Lestidae]]||40||42||29||17||18||18||3||151 |- | [[Perilestidae]]||||19||||||||||||19 |- | [[Synlestidae]]||18||1||8||11||6||||||39 |- | [[Platystictidae]]||136||43||44||||||1||1||224 |- | [[Amphipterygidae]]||||4||||||||||||4 |- | [[Argiolestidae]]||10||||73||19||||||6||108 |- | [[Calopterygidae]]||66||68||5||20||37||8||||185 |- | [[Chlorocyphidae]]||86||||17||42||3||||||144 |- | [[Devadattidae]]||6||||||||||||||6 |- | [[Dicteriadidae]]||||2||||||||||||2 |- | [[Euphaeidae]]||65||||1||||11||||||68 |- | [[Heteragrionidae]]||||51||||||||||||51 |- | [[Hypolestidae]]||||2||||||||||||2 |- | [[Lestoideidae]]||||||9||||||||||9 |- | [[Megapodagrionidae]]||||29||||||||||||29 |- | [[Pentaphlebiidae]]||||||||2||||||||2 |- | [[Philogangidae]]||4||||||||2||||||4 |- | [[Philogeniidae]]||||39||||||||||||39 |- | [[Philosinidae]]||12||||||||||||||12 |- | [[Polythoridae]]||||59||||||||||||59 |- | [[Pseudolestidae]]||1||||||||||||||1 |- | [[Rimanellidae]]||||1||||||||||||1 |- | [[Thaumatoneuridae]]||||5||||||||||||5 |- | ''[[Incertae sedis]]''||25||11||||19||9||||||61 |- | [[Coenagrionidae]]||193||554||152||202||96||103||91||1266 |- | [[Isostictidae]]||||||41||||||||||46 |- | [[Platycnemididae]]||199||||122||70||22||||||404 |- |} Overall, there are about 2942 extant species of damselflies placed in 309 genera.<ref name=diversity>{{cite book |title=Ecology and general biology. Thorp and Covich's Freshwater Invertebrates |edition=4 |publisher=Academic Press |editor=Thorp, James |editor2=D. Christopher Rogers |isbn=978-0-12-385026-3 |author1=Suhling, F. |author2=SahlΓ©n, G. |author3=Gorb, S. |author4=Kalkman, V.J. |author5=Dijkstra, K-D.B. |author6=van Tol, J. |year=2015 |chapter=Order Odonata |pages= 893β932}}</ref>
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