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===Evolution=== {{further |Chimpanzee–human last common ancestor}} Despite a large number of ''[[Homo]]'' fossil finds, ''Pan'' fossils were not described until 2005. Existing chimpanzee populations in West and Central Africa do not overlap with the major [[human fossil]] sites in East Africa, but chimpanzee fossils have now been reported from Kenya. This indicates that both humans and members of the ''Pan'' [[clade]] were present in the [[East African Rift]] Valley during the [[Middle Pleistocene]].<ref name="McBrearty2005">{{cite journal |last1=McBrearty |first1=S. |last2=Jablonski |first2=N. G. |date=September 2005 |title=First fossil chimpanzee |journal=Nature |pmid=16136135 |doi=10.1038/nature04008 |bibcode=2005Natur.437..105M |volume=437 |issue=7055 |pages=105–8|s2cid=4423286}}</ref><ref>{{Cite journal |last=Hopkin |first=Michael |date=2005-08-31 |title=First chimp fossil unearthed |url=https://www.nature.com/news/2005/050829/full/news050829-10.html |journal=Nature |language=en |doi=10.1038/news050829-10 |issn=0028-0836}}</ref><ref>{{Cite web |last=Carey |first=Bjorn |date=2005-08-31 |title=First-ever chimpanzee fossils found |url=https://www.nbcnews.com/id/wbna9145721 |access-date=2025-01-09 |publisher=NBC News |language=en}}</ref> According to studies published in 2017 by researchers at [[George Washington University]], bonobos, along with chimpanzees, split from the human line about 8 million years ago; then bonobos split from the common chimpanzee line about 2 million years ago.<ref name="GWU-20170505">{{cite news |author=Staff |title=Bonobos May Resemble Humans More Than You Think – A GW researcher examined a great ape species' muscles and found they are more closely related to humans than common chimpanzees. |url=https://gwtoday.gwu.edu/bonobos-may-resemble-humans-more-you-think |date=5 May 2017 |publisher=[[George Washington University]] |url-status=live |archiveurl=https://archive.today/20230414135148/https://gwtoday.gwu.edu/bonobos-may-resemble-humans-more-you-think |archivedate=14 April 2023 |accessdate=14 April 2023 }}</ref><ref name="SR-20170404">{{cite journal |last1=Diogo |first1=Rui |last2=Molnar |first2=Julia L. |last3=Wood |first3=Bernard |title=Bonobo anatomy reveals stasis and mosaicism in chimpanzee evolution, and supports bonobos as the most appropriate extant model for the common ancestor of chimpanzees and humans |journal=[[Scientific Reports]] |date=2017 |volume=7 |number=608 |page=608 |doi=10.1038/s41598-017-00548-3 |pmid=28377592 |pmc=5428693 |bibcode=2017NatSR...7..608D }}</ref> Another 2017 genetic study suggests ancient gene flow ([[introgression]]) between 200,000 and 550,000 years ago from the bonobo into the ancestors of central and eastern chimpanzees.<ref name="de Manuel 2016">{{cite journal |last1=de Manuel |first1=M. |first2=M. |last2=Kuhlwilm |last3=P. |first3=Frandsen|display-authors=etal |title=Chimpanzee genomic diversity reveals ancient admixture with bonobos |journal=Science |volume=354 |issue=6311 |pages=477–481 |date=October 2016 |pmid=27789843 |pmc=5546212 |doi=10.1126/science.aag2602 |bibcode=2016Sci...354..477D}}</ref>
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