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== Evolution == [[File:Doedicurus.png|thumb|Illustration of the skeleton of the [[glyptodont]] ''[[Doedicurus]]''.]] The earliest armadillos appeared in South America during the early [[Eocene]] epoch, around 52-55 million years ago.<ref>{{Cite journal |last=Fernicola |first=Juan Carlos |last2=Zimicz |first2=Ana N. |last3=Chornogubsky |first3=Laura |last4=Ducea |first4=Mihai |last5=Cruz |first5=Laura E. |last6=Bond |first6=Mariano |last7=Arnal |first7=Michelle |last8=Cárdenas |first8=Magalí |last9=Fernández |first9=Mercedes |date=September 2021 |title=The Early Eocene Climatic Optimum at the Lower Section of the Lumbrera Formation (Ypresian, Salta Province, Northwestern Argentina): Origin and Early Diversification of the Cingulata |url=https://link.springer.com/10.1007/s10914-021-09545-w |journal=Journal of Mammalian Evolution |language=en |volume=28 |issue=3 |pages=621–633 |doi=10.1007/s10914-021-09545-w |issn=1064-7554}}</ref> The modern groups of armadillos are thought to have diverged from each other in the Eocene.<ref>{{Cite journal |last=Ciancio |first=M. R. |last2=Krmpotic |first2=C. M. |last3=Scarano |first3=A. C. |last4=Epele |first4=M. B. |date=March 2019 |title=Internal Morphology of Osteoderms of Extinct Armadillos and Its Relationship with Environmental Conditions |url=http://link.springer.com/10.1007/s10914-017-9404-y |journal=Journal of Mammalian Evolution |language=en |volume=26 |issue=1 |pages=71–83 |doi=10.1007/s10914-017-9404-y |issn=1064-7554|hdl=11336/41175 |hdl-access=free }}</ref><ref name=":1" /> Recent morphological<ref name=":4">{{Cite journal |last1=Tambusso |first1=P. Sebastián |last2=Varela |first2=Luciano |last3=Góis |first3=Flávio |last4=Moura |first4=Jorge Felipe |last5=Villa |first5=Chiara |last6=Fariña |first6=Richard A. |date=June 2021 |title=The inner ear anatomy of glyptodonts and pampatheres (Xenarthra, Cingulata): Functional and phylogenetic implications |journal=Journal of South American Earth Sciences |language=en |volume=108 |pages=103189 |bibcode=2021JSAES.10803189T |doi=10.1016/j.jsames.2021.103189 |s2cid=234062118}}</ref> and genetic<ref name=":0" /><ref name=":1">{{Cite journal |last=Brambilla |first=Luciano |last2=Ibarra |first2=Damián A. |last3=Barboza |first3=María C. |last4=Bresso |first4=Edgardo G. |last5=Rosano |first5=Germán |last6=Pérez |first6=Germán |last7=Straccia |first7=Pablo |last8=Scian |first8=Rubén D. |last9=Brun |first9=Lucas R. |date=February 2025 |title=Mitochondrial genome of Neuryurus rudis (Xenarthra, Cingulata); contribution to phylogeny and origin of glyptodonts |url=https://linkinghub.elsevier.com/retrieve/pii/S0378111924009405 |journal=Gene |language=en |volume=936 |pages=149059 |doi=10.1016/j.gene.2024.149059}}</ref> evidence indicates that the large [[Glyptodont|glyptodonts]] and [[Pampatheriidae|pampatheres]], both of which went extinct as part of the [[Late Pleistocene extinctions|end-Pleistocene extinction event]] around 12,000 years ago, are nested within the diversity of living armadillos. Below is a recent simplified phylogeny of the [[Cingulata]] based on genetics and analysis of the inner ear.<ref name=":4" /> The dagger symbol, "†", denotes extinct groups.{{clade|{{Clade |1= [[Dasypodidae]] |2={{clade |1=[[Euphractinae]] |2={{clade |1=[[Chlamyphorinae]] |2=[[Tolypeutinae]] |3={{clade |1={{extinct}}[[Pampatheres]] |2={{extinct}}[[Glyptodonts]] }}}}}}}}|label1=[[Cingulata]]}}
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