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===Natural predators=== [[File:20130825_Brazil_Rio_de_Janeiro_0144.jpg|thumb|left|[[Theropod]] dinosaur ''[[Irritator]]'' shown feeding on a pterosaur]] Pterosaurs are known to have been eaten by [[theropoda|theropods]]. In the 1 July 2004 edition of ''[[Nature (journal)|Nature]]'', paleontologist [[Éric Buffetaut]] discusses an Early Cretaceous fossil of three cervical [[vertebra]]e of a pterosaur with the broken tooth of a [[spinosaur]], most likely ''[[Irritator]]'', embedded in it. The vertebrae are known not to have been eaten and exposed to digestion, as the joints are still articulated.<ref>{{cite journal |vauthors=Buffetaut E, Martill D, Escuillié F |title=Pterosaurs as part of a spinosaur diet |journal=Nature |volume=430 |issue=6995 |page=33 |date=July 2004 |pmid=15229562 |doi=10.1038/430033a |bibcode=2004Natur.429...33B|s2cid=4398855 |doi-access=free }}</ref> Fossils of ''Pteranodon'' have been found with tooth marks from sharks such as ''[[Squalicorax]]'',<ref>{{Cite web|url=https://www.nationalgeographic.com/science/2018/10/news-sharks-eating-pterosaurs-fossils-cretaceous-paleontology/|archive-url=https://web.archive.org/web/20181003163916/https://www.nationalgeographic.com/science/2018/10/news-sharks-eating-pterosaurs-fossils-cretaceous-paleontology/|url-status=dead|archive-date=October 3, 2018|title=Prehistoric sharks feasted on flying reptiles, fossil reveals|date=October 3, 2018|website=Science & Innovation}}</ref> and a fossil with tooth marks from the ''[[Toolebuc formation]]'' has been interpreted as being attacked or scavenged by an [[ichthyosaur]] (most likely ''[[Platypterygius]]'').
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