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== Properties for animal nutrition == Tannins have traditionally been considered [[Antinutrient|antinutritional]], depending upon their chemical structure and dosage.<ref>{{cite journal |author1=Muller-Harvey I. |author2=McAllan A. B. |title=Tannins: Their biochemistry and nutritional properties |journal=Advances in Plant Cell Biochemistry and Biotechnology |volume=1 |pages=151β217 |year=1992}}</ref> Many studies suggest that chestnut tannins have positive effects on [[silage]] quality in the [[Baler#Silage or haylage bales|round bale silages]], in particular reducing [[Non-protein nitrogen|NPN]]s (non-protein nitrogen) in the lowest wilting level.<ref>{{cite journal |author1=Tabacco E. |author2=Borreani G. |author3=Crovetto G. M. |author4=Galassi G. |author5=Colombo D. |author6=Cavallarin L. |title=Effect of chestnut tannin on fermentation quality, proteolysis, and protein rumen degradability of alfalfa silage |journal=Journal of Dairy Science |volume=89 |issue=12 |pages=4736β4746 |date=1 December 2006 |pmid=17106105 |doi=10.3168/jds.S0022-0302(06)72523-1 |doi-access=free |df=dmy-all }}</ref> Improved fermentability of [[Soybean|soya meal]] nitrogen in the [[rumen]] may occur.<ref>{{cite journal |doi=10.1051/animres:19930210 |author1=Mathieu F. |author2=Jouany J. P. |title=Effect of chestnut tannin on the fermentability of soyabean meal nitrogen in the rumen |journal=Ann Zootech |volume=42 |page=127 |year=1993 |issue=2 |doi-access=free }}</ref> Condensed tannins inhibit herbivore digestion by binding to consumed plant proteins and making them more difficult for animals to digest, and by interfering with protein absorption and digestive enzymes (for more on that topic, see [[plant defense against herbivory]]). [[Histatin]]s, another type of [[salivary protein]]s, also precipitate tannins from solution, thus preventing alimentary adsorption.<ref>{{cite journal | last=Shimada | first=Takuya | title=Salivary Proteins as a Defense Against Dietary Tannins | journal=[[Journal of Chemical Ecology]] | volume=32 | issue=6 | date=23 May 2006 | pmid=16770710 | doi=10.1007/s10886-006-9077-0 | pages=1149β1163| bibcode=2006JCEco..32.1149S | s2cid=21617545 }}</ref> [[Legume]] fodders containing condensed tannins are a possible option for integrated sustainable control of gastrointestinal [[nematode]]s in ruminants, which may help address the worldwide development of resistance to synthetic [[anthelmintics]]. These include nuts, temperate and tropical barks, carob, coffee and cocoa.<ref name="Hoste2022">{{cite journal | last1=Hoste | first1=HervΓ© | last2=Meza-OCampos | first2=Griselda | last3=Marchand | first3=Sarah | last4=Sotiraki | first4=Smaragda | last5=Sarasti | first5=Katerina | last6=Blomstrand | first6=Berit M. | last7=Williams | first7=Andrew R. | last8=Thamsborg | first8=Stig M. | last9=Athanasiadou | first9=Spiridoula | last10=Enemark | first10=Heidi L. | last11=Torres Acosta | first11=Juan Felipe | last12=Mancilla-Montelongo | first12=Gabriella | last13=Castro | first13=Carlos Sandoval | last14=Costa-Junior | first14=Livio M. | last15=Louvandini | first15=Helder | last16=Sousa | first16=Dauana Mesquita | last17=Salminen | first17=Juha-Pekka | last18=Karonen | first18=Maarit | last19=Engstrom | first19=Marika | last20=Charlier | first20=Johannes | last21=Niderkorn | first21=Vincent | last22=Morgan | first22=Eric R. | title=Use of agro-industrial by-products containing tannins for the integrated control of gastrointestinal nematodes in ruminants | journal=Parasite | volume=29 | year=2022 | doi=10.1051/parasite/2022010 | page=10| pmid=35225785 | pmc=8884022 }} {{open access}}</ref>
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