Editing Respiratory system (section)

====Insects====
{{Main|Respiratory system of insects}}
Most insects breath passively through their [[Spiracle (arthropods)|spiracle]]s (special openings in the [[exoskeleton]]) and the air reaches every part of the body by means of a series of smaller and smaller tubes called 'trachaea' when their diameters are relatively large, and '[[tracheole]]s' when their diameters are very small. The tracheoles make contact with individual cells throughout the body.<ref name=campbell /> They are partially filled with fluid, which can be withdrawn from the individual tracheoles when the tissues, such as muscles,  are active and have a high demand for oxygen, bringing the air closer to the active cells.<ref name=campbell /> This is probably brought about by the buildup of lactic acid in the active muscles causing an osmotic gradient, moving the water out of the tracheoles and into the active cells.  Diffusion of gases is effective over small distances but not over larger ones, this is one of the reasons insects are all relatively small. Insects which do not have spiracles and trachaea, such as some Collembola, breathe directly through their skins, also by diffusion of gases.<ref>[http://www.earthlife.net/insects/anatomy.html The Earth Life Web, Insect Morphology and Anatomy]. Earthlife.net. Retrieved on 2013-04-21.</ref>

The number of spiracles an insect has is variable between species, however, they always come in pairs, one on each side of the body, and usually one pair per segment. Some of the Diplura have eleven, with four pairs on the thorax, but in most of the ancient forms of insects, such as Dragonflies and Grasshoppers there are two thoracic and eight abdominal spiracles. However, in most of the remaining insects, there are fewer. It is at the level of the tracheoles that oxygen is delivered to the cells for respiration.

Insects were once believed to exchange gases with the environment continuously by the [[simple diffusion]] of gases into the tracheal system. More recently, however, large variation in insect ventilatory patterns has been documented and insect respiration appears to be highly variable. Some small insects do not demonstrate continuous respiratory movements and may lack muscular control of the spiracles. Others, however, utilize [[muscle contraction|muscular contraction]] of the [[abdomen]] along with coordinated spiracle contraction and relaxation to generate cyclical gas exchange patterns and to reduce water loss into the atmosphere. The most extreme form of these patterns is termed [[discontinuous gas exchange]] cycles.<ref>{{cite journal
 | last = Lighton
 | first = JRB
 | title = Discontinuous gas exchange in insects
 | journal = Annu Rev Entomol
 | volume = 41
 | pages = 309–324
 | date = January 1996
 | doi = 10.1146/annurev.en.41.010196.001521 | pmid = 8546448
 }}</ref>