Editing Pterosaur (section)

===Swimming===
Tracks made by ctenochasmatoids indicate that these pterosaurs swam using their hindlimbs. In general, these have large hindfeet and long torsos, indicating that they were probably more adapted for swimming than other pterosaurs.<ref name= "witton2013">{{harvnb|Witton|2013|p=51}}</ref> Pteranodontians conversely have several speciations in their humeri interpreted to have been suggestive of a water-based version of the typical quadrupedal launch, and several like [[boreopterid]]s must have foraged while swimming, as they seem incapable of [[frigatebird]]-like aerial hawking.<ref name="witton2013"/> These adaptations are also seen in terrestrial pterosaurs like [[azhdarchid]]s, which presumably still needed to launch from water in case they found themselves in it. The [[nyctosaurid]] ''[[Alcione (pterosaur)|Alcione]]'' may display adaptations for wing-propelled diving like modern [[gannet]]s and [[tropicbird]]s.<ref name="longrichetal2018" />