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=== Terrestrial mammals === {{further|Shrews#Echolocation}} Terrestrial mammals other than bats known or thought to echolocate include [[shrew]]s,<ref>{{cite journal |last1=Tomasi |first1=Thomas E. |year=1979 |title=Echolocation by the Short-Tailed Shrew ''Blarina brevicauda'' | journal=Journal of Mammalogy |volume=60 |issue=4 | pages=751β759 |doi=10.2307/1380190 | jstor=1380190 }}</ref><ref>{{Cite journal |last=Buchler |first=E. R. |date=November 1976 |title=The use of echolocation by the wandering shrew (Sorex vagrans) |url=https://linkinghub.elsevier.com/retrieve/pii/S0003347276800164 |journal=Animal Behaviour |volume=24 |issue=4 |pages=858β873 |doi=10.1016/S0003-3472(76)80016-4 |s2cid=53160608 }}</ref><ref name="Siemers">{{cite journal |last1=Siemers |first1=BjΓΆrn M. |last2=Schauermann |first2=Grit |last3=Turni |first3=Hendrik |last4=von Merten |first4=Sophie |title=Why do shrews twitter? Communication or simple echo-based orientation |journal=Biology Letters |volume=5 |issue=5 |pages=593β596 |date=October 2009 |pmid=19535367 |pmc=2781971 |doi=10.1098/rsbl.2009.0378 }}</ref> the [[tenrec]]s of [[Madagascar]],<ref>{{cite journal |last=Gould |first=Edwin |title=Evidence for echolocation in the Tenrecidae of Madagascar |journal=Proceedings of the American Philosophical Society |volume=109 |issue=6 |year=1965 |pages=352β360 |jstor=986137 }}</ref> [[Chinese pygmy dormouse|Chinese pygmy dormice]],<ref>{{cite journal |last1=He |first1=Kai |last2=Liu |first2=Qi |last3=Xu |first3=Dong-Ming |last4=Qi |first4=Fei-Yan |last5=Bai |first5=Jing |last6=He |first6=Shui-Wang |last7=Chen |first7=Peng |last8=Zhou |first8=Xin |last9=Cai |first9=Wan-Zhi |last10=Chen |first10=Zhong-Zheng |last11=Liu |first11=Zhen |last12=Jiang |first12=Xue-Long |last13=Shi |first13=Peng |display-authors=3 |date=2021-06-18 |title=Echolocation in soft-furred tree mice |url=https://www.science.org/doi/10.1126/science.aay1513 |journal=Science |volume=372 |issue=6548 |pages=eaay1513 |doi=10.1126/science.aay1513 |pmid=34140356 |s2cid=235463083 |issn=0036-8075}}</ref> and [[solenodon]]s.<ref name="Eisenberg1966">{{cite journal |last1=Eisenberg |first1=J. F. |last2=Gould |first2=E. |year=1966 |title=The Behavior of ''Solenodon paradoxus'' in Captivity with Comments on the Behavior of other Insectivora |journal=[[Zoologica]] |volume=51 |issue=4 |pages=49β60 |df=dmy-all }}</ref> Shrew sounds, unlike those of bats, are low amplitude, broadband, multi-harmonic and frequency modulated.<ref name="Siemers"/> They contain no echolocation clicks with reverberations, and appear to be used for simple, close range spatial orientation. In contrast to bats, shrews use echolocation only to investigate their habitat rather than to pinpoint food.<ref name="Siemers"/> There is evidence that blinded [[laboratory rats]] can use echolocation to navigate mazes.<ref>{{cite journal |last1=Riley |first1=Donald A. |last2=Rosenzweig |first2=Mark R. |title=Echolocation in Rats |journal=Journal of Comparative and Physiological Psychology |volume=50 |issue=4 |pages=323β328 |date=August 1957 |pmid=13475510 |doi=10.1037/h0047398 }}</ref>
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