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=== Innervation === The glans and the frenulum are [[nerve|innervated]] by the bilateral [[dorsal nerve of the penis]] and the [[perineal nerve]], both divisions of the [[pudendal nerve]].<ref name=":6">{{Citation |last1=Weech |first1=David |title=Anatomy, Abdomen and Pelvis, Penis Dorsal Nerve |date=2022 |url=http://www.ncbi.nlm.nih.gov/books/NBK525966/ |work=StatPearls |place=Treasure Island (FL) |publisher=StatPearls Publishing |pmid=30247841 |access-date=2022-09-26 |last2=Ameer |first2=Muhammad Atif |last3=Ashurst |first3=John V. |archive-date=2023-04-05 |archive-url=https://web.archive.org/web/20230405190749/https://www.ncbi.nlm.nih.gov/books/NBK525966/ |url-status=live }}</ref> Branches of the dorsal nerve extend through the glans ventrolaterally displaying a three-dimensional innervation pattern.<ref name=":6"/><ref name=":2">{{Cite journal |last1=Yang |first1=Claire C. |last2=Bradley |first2=William E. |date=1999-01-01 |title=Innervation of the human glans penis |url=https://www.auajournals.org/doi/abs/10.1016/S0022-5347(01)62075-5 |journal=Journal of Urology |volume=161 |issue=1 |pages=97β102 |doi=10.1016/S0022-5347(01)62075-5 |pmid=10037378 |access-date=2022-09-24 |archive-date=2021-11-21 |archive-url=https://web.archive.org/web/20211121191429/https://www.auajournals.org/doi/abs/10.1016/S0022-5347%2801%2962075-5 |url-status=live }}</ref> The main branches form smaller bundles of nerves that expand outwards into the tissue of the glans.<ref name=":6"/> The rich innervation of the glans penis reveals its function as a primary anatomical source of male sexual pleasure.<ref name="yangcc"/><ref name=":2"/><ref>{{Cite book |last1=Greenberg |first1=Jerrold |url=https://books.google.com/books?id=1NC5R0RozBYC&pg=PP1 |title=Exploring the Dimensions of Human Sexuality |last2=Bruess |first2=Clint |last3=Conklin |first3=Sarah |date=2010-03-10 |publisher=Jones & Bartlett Learning |isbn=978-0-7637-7660-2 |pages=132 |language=en}}</ref> Yang & Bradley argue; "the distinct pattern of innervation of the glans emphasizes its role as a sensory structure".<ref name="yangcc"/> While Yang & Bradley's (1998) report "showed no areas in the glans to be more densely innervated than others.",<ref name="yangcc">{{cite journal | last = Yang | first = C. C. |author2=W.E. Bradley |date=July 1998 | title = Neuroanatomy of the penile portion of the human dorsal nerve of the penis | journal = British Journal of Urology | volume = 82 | issue = 1 | pages = 109β13 | doi =10.1046/j.1464-410x.1998.00669.x | pmid = 9698671 }}</ref> Halata & Munger (1986) report that the density of several nerve terminals is greatest in the corona glandis.<ref name=":5"/> Halata & Spathe (1997) reported: "The glans penis contains a predominance of [[free nerve ending]]s, numerous genital [[bulboid corpuscles|end bulbs]] and rarely [[Pacinian corpuscle|Pacinian]] and [[Bulbous corpuscle|Ruffinian]] corpuscles. [[Merkel nerve ending]]s and [[Meissner's corpuscle]]s ([[mechanoreceptor]]s typically found in thick [[glabrous skin]]) are not present".<ref name="halata2">{{Cite book | last = Halata | first = Zdenek | author2 = A. Spaethe | chapter = Sensory Innervation of the Human Penis | year = 1997 | title = The Fate of the Male Germ Cell | pages = 265β6 | pmid = 9361804 | url = http://www.cirp.org/library/anatomy/halata2/ | access-date = 2006-07-07 | volume = 424 | doi = 10.1007/978-1-4615-5913-9_48 | series = Advances in Experimental Medicine and Biology | isbn = 978-0-306-45696-1 | archive-date = 2006-06-20 | archive-url = https://web.archive.org/web/20060620015859/http://www.cirp.org/library/anatomy/halata2/ | url-status = live }}</ref> The genital end bulbs, that are present throughout the glans, are most numerous in the corona and near the frenulum.<ref name=":5">{{Cite journal |last1=Halata |first1=Z. |last2=Munger |first2=B. L. |date=1986-04-23 |title=The neuroanatomical basis for the protopathic sensibility of the human glans penis |url=https://pubmed.ncbi.nlm.nih.gov/3697758/ |journal=Brain Research |volume=371 |issue=2 |pages=205β230 |doi=10.1016/0006-8993(86)90357-4 |issn=0006-8993 |pmid=3697758 |s2cid=23781274 |access-date=2022-09-25 |archive-date=2022-09-25 |archive-url=https://web.archive.org/web/20220925152126/https://pubmed.ncbi.nlm.nih.gov/3697758/ |url-status=live }}</ref> Simple, Pacinian and Ruffinian corpuscles are identified predominantly in the corona glandis. The most numerous nerve terminals are free nerve endings present in almost every [[dermal papilla]] of the glans, as well as scattered throughout the deeper [[dermis]].<ref name=":5"/>
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