Editing Clouded leopard (section)

=== Phylogeny ===
Skulls of clouded leopard and ''[[Panthera]]'' species were analysed [[Morphology (biology)|morphologically]] in the 1960s. Results indicate that the clouded leopard forms an [[evolution]]ary link between the [[Pantherinae]] and the [[Felinae]].<ref name=Hemmer1968 />
[[Phylogenetic]] analysis of the [[nuclear DNA]] in tissue samples from all Felidae species revealed that the [[evolutionary radiation]] of the Felidae began in the [[Miocene]] around {{mya|14.45|8.38}} in Asia.<ref name=Johnson2006>{{cite journal |last1=Johnson |first1=W. E. |last2=Eizirik |first2=E. |last3=Pecon-Slattery |first3=J. |last4=Murphy, W. J. |last5=Antunes, A. |last6=Teeling, E. |last7=O'Brien, S. J. |name-list-style=amp |year=2006 |doi=10.1126/science.1122277 |title=The Late Miocene radiation of modern Felidae: A genetic assessment |journal=[[Science (journal)|Science]] |volume=311 |pages=73–77 |issue=5757 |url=https://zenodo.org/record/1230866 |pmid=16400146 |bibcode=2006Sci...311...73J |s2cid=41672825 |access-date=11 September 2019 |archive-date=4 October 2020 |archive-url=https://web.archive.org/web/20201004075725/https://zenodo.org/record/1230866 |url-status=live}}</ref><ref name=Werdelin2010>{{cite book |last1=Werdelin |first1=L. |last2=Yamaguchi |first2=N. |last3=Johnson |first3=W. E. |last4=O'Brien |first4=S. J. |name-list-style=amp |chapter=Phylogeny and evolution of cats (Felidae) |year=2010 |pages=59–82 |editor1-last=Macdonald |editor1-first=D. W. |editor2-last=Loveridge |editor2-first=A. J. |title=Biology and Conservation of Wild Felids |publisher=Oxford University Press |location=Oxford, UK |isbn=978-0-19-923445-5 |chapter-url=https://www.researchgate.net/publication/266755142 |access-date=16 September 2019 |archive-date=25 September 2018 |archive-url=https://web.archive.org/web/20180925141956/https://www.researchgate.net/publication/266755142 |url-status=live}}</ref> Analysis of [[mitochondrial DNA]] of all Felidae species indicates a radiation at {{mya|16.76|6.46}}.<ref name=Li_al2016>{{cite journal |author=Li, G. |author2=Davis, B. W. |author3=Eizirik, E. |name-list-style=amp |author4=Murphy, W. J. |year=2016 |title=Phylogenomic evidence for ancient hybridization in the genomes of living cats (Felidae) |journal=Genome Research |volume=26 |issue=1 |pages=1–11 |doi=10.1101/gr.186668.114 |pmid=26518481 |pmc=4691742}}</ref>
The clouded leopard is the [[Sister group|sister taxon]] to all other members of the Pantherinae, diverging {{mya|9.32|4.47}}, based on analysis of their nuclear DNA.<ref name=Johnson2006/>
The clouded leopard from mainland Asia reached [[Borneo]] and [[Sumatra]] via a now submerged [[land bridge]] probably during the [[Pleistocene]], when populations became isolated during periods of global cooling and warming. [[Genetic analysis]] of hair samples of the clouded leopard and its [[Sister group|sister species]] the Sunda clouded leopard (''N. diardi'') indicates that they diverged 2.0–0.93 million years ago.<ref name=bb/>

{{clade gallery |align=left |width=400px; |main-caption=Phylogenetic relationships of the clouded leopard as derived through analysis of
|header1=nuclear DNA:<ref name=Johnson2006/>
|cladogram1={{clade |label1=Felidae
  |1={{clade |label2=Pantherinae
   |1=Felinae
   |2={{clade |1=''Panthera'' |2=Clouded leopard }} }} }}
|header2=combined mitochondrial and nuclear DNA:<ref name=bb/>
|cladogram2={{clade |label1=Felidae
  |1={{clade |label2=Pantherinae
   |1=Felinae
   |2={{clade
    |1=''Panthera'' |label2=''Neofelis''
    |2={{clade
     |1=Sunda clouded leopard
     |2=Mainland clouded leopard }} }} }} }} }}
{{clear|left}}