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====Smell==== [[File:EB1911 Chiroptera Fig. 5.jpg|thumb|alt=A scientific illustration of a megabat face in profile with prominent nostrils. Each nostril is a distinct tube projecting away from the face at a right angle.|The nostrils of the [[island tube-nosed fruit bat]] (''Nyctimene major'')]] Megabats use smell to find food sources like fruit and nectar.<ref name="Jones"/> They have keen senses of smell that rival that of the [[domestic dog]].<ref name="Schwab">{{cite journal|doi=10.1136/bjo.2005.077966|pmid=16267906|pmc=1772916|title=A choroidal sleight of hand|journal=[[British Journal of Ophthalmology]]|volume=89|issue=11|page=1398|year=2005|last1=Schwab|first1=I. R.}}</ref> Tube-nosed fruit bats such as the [[eastern tube-nosed bat]] (''Nyctimene robinsoni'') have stereo [[olfaction]], meaning they are able to map and follow odor plumes three-dimensionally.<ref name="Schwab"/> Along with most (or perhaps all) other bat species, megabats mothers and offspring also use scent to recognize each other, as well as for recognition of individuals.<ref name="Jones">{{cite journal|doi=10.3389/fphys.2013.00117|pmid=23755015|pmc=3667242|title=From the ultrasonic to the infrared: Molecular evolution and the sensory biology of bats|journal=[[Frontiers in Physiology]]|volume=4|page=117|year=2013|last1=Jones|first1=G.|last2=Teeling|first2=E. C.|last3=Rossiter|first3=S. J.|doi-access=free}}</ref> In flying foxes, males have enlarged [[androgen]]-sensitive [[sebaceous gland]]s on their shoulders they use for [[Territory (animal)#Spraying|scent-marking]] their territories, particularly during the mating season. The secretions of these glands vary by species—of the 65 chemical compounds isolated from the glands of four species, no compound was found in all species.<ref name="Wood 2005"/> Males also engage in [[Urination#Other animals|urine washing]], or coating themselves in their own urine.<ref name="Wood 2005">{{cite journal| last1=Wood| first1= W. F.| last2= Walsh| first2= A.| last3= Seyjagat| first3= J.| last4= Weldon| first4= P. J.| title= Volatile Compounds in Shoulder Gland Secretions of Male Flying Foxes, Genus Pteropus (Pteropodidae, Chiroptera)| year=2005| journal=[[Z Naturforsch C]]| volume=60| issue=9–10| pages= 779–784|url=https://www.researchgate.net/publication/7447695| doi= 10.1515/znc-2005-9-1019| pmid= 16320623| s2cid= 1938648| doi-access= free}}</ref><ref name="Wagner 2008">{{cite journal| last=Wagner| first= J.| year=2008| title= Glandular secretions of male Pteropus (Flying foxes): preliminary chemical comparisons among species|journal=Independent Study Project (Isp) Collection| url=http://digitalcollections.sit.edu/isp_collection/559/}}</ref>
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