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===Hunting and feeding=== {{main article|Hunting behavior of gray wolves}} [[File:Wolves and elk (cropped).jpg|thumb|right|alt=Aerial photograph a bull elk in winter being pursued by four wolves |Wolves pursuing a bull elk]] Single wolves or mated pairs typically have higher success rates in hunting than do large packs; single wolves have occasionally been observed to kill large prey such as moose, [[bison]] and [[muskox]]en unaided.{{sfn|Mech|Boitani|2003|pp=119β121}}<ref name=Thurber1993/> The size of a wolf hunting pack is related to the number of pups that survived the previous winter, adult survival, and the rate of dispersing wolves leaving the pack. The optimal pack size for hunting elk is four wolves, and for bison a large pack size is more successful.{{sfn|Mech|Smith|MacNulty|2015|p=4}} Wolves move around their territory when hunting, using the same trails for extended periods.{{sfn|Heptner|Naumov|1998|p=233}} Wolves are nocturnal predators. During the winter, a pack will commence hunting in the twilight of early evening and will hunt all night, traveling tens of kilometres. Sometimes hunting large prey occurs during the day. During the summer, wolves generally tend to hunt individually, ambushing their prey and rarely giving pursuit.{{sfn|Heptner|Naumov|1998|p=240}} When hunting large gregarious prey, wolves will try to isolate an individual from its group.<ref name=MacNulty2007/> If successful, a wolf pack can bring down game that will feed it for days, but one error in judgement can lead to serious injury or death. Most large prey have developed defensive adaptations and behaviours. Wolves have been killed while attempting to bring down bison, elk, moose, muskoxen, and even by one of their smallest hoofed prey, the white-tailed deer. With smaller prey like [[beaver]], geese, and hares, there is no risk to the wolf.{{sfn|Mech|Smith|MacNulty|2015|pp=1β3}} Although people often believe wolves can easily overcome any of their prey, their success rate in hunting hoofed prey is usually low.{{sfn|Mech|Smith|MacNulty|2015|p=7}} [[File:Wolves eating deer.jpeg|thumb|right|alt=Photograph of two wolves eating a deer carcass at night|Two wolves feeding on a [[white-tailed deer]]]] The wolf must give chase and gain on its fleeing prey, slow it down by biting through thick hair and hide, and then disable it enough to begin feeding.{{sfn|Mech|Smith|MacNulty|2015|pp=1β3}} Wolves may wound large prey and then lie around resting for hours before killing it when it is weaker due to blood loss, thereby lessening the risk of injury to themselves.{{sfn|Mech|Smith|MacNulty|2015|p=82β89}} With medium-sized prey, such as roe deer or [[sheep]], wolves kill by [[Throat clamp|biting the throat]], severing nerve tracks and the [[carotid artery]], thus causing the animal to die within a few seconds to a minute. With small, [[mouse]]like prey, wolves leap in a high arc and immobilize it with their forepaws.<ref name=Zimen/> Once prey is brought down, wolves begin to feed excitedly, ripping and tugging at the carcass in all directions, and bolting down large chunks of it.{{sfn|Mech|1981|p=185}} The breeding pair typically monopolizes food to continue producing pups. When food is scarce, this is done at the expense of other family members, especially non-pups.{{sfn|Mech|Boitani|2003|p=58}} Wolves typically commence feeding by gorging on the larger internal organs, like the [[heart]], [[liver]], [[lung]]s, and [[stomach]] lining. The [[kidney]]s and [[spleen]] are eaten once they are exposed, followed by the muscles.{{sfn|Mech|Boitani|2003|pp=122β125}} A wolf can eat 15β19% of its body weight in one sitting.{{sfn|Mech|Boitani|2003|p=201}}
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