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=== Holism === {{Main|Holism}} Holism remains a critical part of the theoretical foundation in contemporary ecological studies. Holism addresses the [[Biological organisation|biological organization]] of life that [[Systems biology|self-organizes]] into layers of emergent whole systems that function according to non-reducible properties. This means that higher-order patterns of a whole functional system, such as an [[ecosystem]], cannot be predicted or understood by a simple summation of the parts.<ref name="Liu09"/> "New properties emerge because the components interact, not because the basic nature of the components is changed."<ref name="Odum05"/>{{rp|8}} Ecological studies are necessarily holistic as opposed to [[reductionistic]].<ref name="Levins80" /><ref name="Odum1977"/><ref name="Mikkelson10"/> Holism has three scientific meanings or uses that identify with ecology: 1) the mechanistic complexity of ecosystems, 2) the practical description of patterns in quantitative reductionist terms where correlations may be identified but nothing is understood about the causal relations without reference to the whole system, which leads to 3) a [[metaphysics|metaphysical]] hierarchy whereby the causal relations of larger systems are understood without reference to the smaller parts. Scientific holism differs from [[mysticism]] that has appropriated the same term. An example of metaphysical holism is identified in the trend of increased exterior thickness in shells of different species. The reason for a thickness increase can be understood through reference to principles of natural selection via predation without the need to reference or understand the [[biomolecular]] properties of the exterior shells.<ref name="Wilson88"/>
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