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==Other animals== {{Main|Animal sexual behavior|Non-reproductive sexual behavior in animals#Oral sex}} [[File:Cynopterus sphinx fellatio no music.ogg|thumb|Female bats perform fellatio to increase copulation time. This species is the only non-primate known to exhibit this behaviour.<ref name=plos />]] [[Pteropus|Flying foxes]] (a type of bat) have been observed engaging in oral sex.<ref name="Sugita 2016">Sugita, N. (2016). Homosexual Fellatio: Erect Penis Licking between Male Bonin Flying Foxes Pteropus pselaphon. PLoS ONE, 11(11), e0166024.</ref> [[Indian flying fox]] males will lick a female's vulva both before and after [[copulation (zoology)|copulation]], with the length of pre-copulation [[cunnilingus]] positively correlated with length of copulation.<ref name="Maruthupandian 2013">Maruthupandian, J., & Marimuthu, G. (2013). Cunnilingus apparently increases duration of copulation in the Indian flying fox, Pteropus giganteus. PLoS ONE, 8(3), e59743.</ref> The [[fruit bat]] ''[[Cynopterus sphinx]]'', has been observed to engage in fellatio during mating. Pairs spend more time copulating if the female licks the male than if she does not.<ref name="plos">{{cite journal|last=Tan|first=Min|author2=Gareth Jones |author3=Guangjian Zhu |author4=Jianping Ye |author5=Tiyu Hong |author6=Shanyi Zhou |author7=Shuyi Zhang |author8=Libiao Zhang |date=October 28, 2009|title=Fellatio by Fruit Bats Prolongs Copulation Time|journal=[[PLoS ONE]]|volume=4|issue=10|doi=10.1371/journal.pone.0007595|pmid=19862320|pages=e7595|pmc=2762080|bibcode=2009PLoSO...4.7595T|editor1-last=Hosken|editor1-first=David|doi-access=free}}</ref><ref>{{cite web |url=https://www.newscientist.com/article/dn18077-fellatio-keeps-male-fruit-bats-keen.html |title=Fellatio keeps male fruit bats keen - life - 29 October 2009 - New Scientist |publisher=www.newscientist.com |access-date=2009-10-31 |archive-date=2015-04-26 |archive-url=https://web.archive.org/web/20150426103655/http://www.newscientist.com/article/dn18077-fellatio-keeps-male-fruit-bats-keen.html |url-status=live }}</ref> Male [[Livingstone's fruit bat]]s have been observed engaging in [[Homosexual behavior in animals|homosexual]] fellatio, although it is unknown if this is an example of sexual behavior or [[social grooming]].<ref name="Courts 1996">Courts, S. E. (1996). An ethogram of captive Livingstone's fruit bats Pteropus livingstonii in a new enclosure at Jersey Wildlife Preservation Trust. The Dodo: Journal of the Wildlife Preservation Trusts, 32, 15β37.</ref> [[Bonin flying fox]]es also engage in homosexual fellatio, but the behavior has been observed independently of social grooming.<ref name="Sugita 2016"/>
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