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=== Pests and predators === [[File:Scarlet lily beetle lilioceris lilii.jpg|thumb|left|upright=0.8|[[Scarlet lily beetle]] (''Lilioceris lilii'')]] [[File:Semper Augustus Tulip 17th century.jpg|thumb|left|upright=0.8|17th-century painting of affected ''Tulipa'' 'Semper Augustus', showing '[[tulip breaking virus|breaking]]']] Liliaceae are subject to a wide variety of diseases and pests, including [[insects]], such as [[thrips]], [[aphids]], [[beetles]] and [[flies]]. Also [[fungi]], [[viruses]] and [[vertebrate]] animals such as [[mice]] and [[deer]].<ref name=UCIPML/><ref name=UCIPMT/> An important horticultural and garden pest is the [[scarlet lily beetle]] (Japanese red lily beetle, ''Lilioceris lilii'') and other [[Lilioceris]] species which attack ''[[Fritillaria]]'' and ''[[Lilium]]''.<ref name="Majka"/> ''Lilium'' species may be food plants for the [[Cosmia trapezina]] moth. A major pest of Tulips is the fungus, ''[[Botryotinia|Botrytis tulipae]]''. Both ''Lilium'' and ''Tulipa'' are susceptible to a group of five viruses of the family ''[[Potyviridae]]'', specifically the [[potyvirus]] (named for ''[[potato virus Y]])'' group, which includes the ''[[tulip-breaking virus]]'' (''TBV'') and the ''lily streak virus'' (lily mottle virus, ''LMoV'') resulting in 'breaking' of the color of the flowers. The viruses are transmitted by [[aphids]]. This breaking effect was of economic importance during the [[tulip mania]] of the seventeenth century, because it appeared to be producing new varieties.<ref name=Phillips/><ref name=Garber/> In modern times tulip breeders have produced varieties that mimic the effect of the virus, without being infected. One of these varieties is known as 'Rembrandt', after the [[Rembrandt|Dutch artist of that name]]. Contemporary tulip owners commonly had Rembrandt and other artists paint their flowers to preserve them for posterity, hence the 'broken' tulips were known as Rembrandt tulips at that time. Another modern variety is 'Princess Irene'.<ref name=Beckerman/><ref name="Esther Inglis-Arkell"/><ref name=RPD634/> One of the tulip breaking viruses is also named the ''Rembrandt tulip-breaking virus'' (ReTBV).<ref name=Dekker/><ref name=UniprotReTBV/>
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