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=== Hologenome development and evolution === Evolution originated from changes in development where variations within species are selected for or against because of the symbionts involved.<ref name="Li-2015">{{cite journal |last1=Li |first1=Ci-Xiu |last2=Shi |first2=Mang |last3=Tian |first3=Jun-Hua |last4=Lin |first4=Xian-Dan |last5=Kang |first5=Yan-Jun |last6=Chen |first6=Liang-Jun |last7=Qin |first7=Xin-Cheng |last8=Xu |first8=Jianguo |last9=Holmes |first9=Edward C |last10=Zhang |first10=Yong-Zhen |display-authors=3 |title=Unprecedented genomic diversity of RNA viruses in arthropods reveals the ancestry of negative-sense RNA viruses |journal=eLife |volume=4 |issue=|pages=|date=January 2015 |pmid=25633976 |pmc=4384744 |doi=10.7554/eLife.05378 |url=|doi-access=free }}</ref> The hologenome theory relates to the holobiont and symbionts genome together as a whole.<ref>{{cite journal |last1=Rosenberg |first1=E. |last2=Zilber-Rosenberg |first2=I. |title=Symbiosis and development: the hologenome concept |journal=Birth Defects Research. Part C, Embryo Today |volume=93 |issue=1 |pages=56β66 |date=March 2011 |pmid=21425442 |doi=10.1002/bdrc.20196 }}</ref> Microbes live everywhere in and on every multicellular organism.<ref>{{cite journal |last=Morris |first=J.J. |title=What is the hologenome concept of evolution? |journal=F1000Research |volume=7 |pages=1664 |date=2018-10-19 |pmid=30410727 |pmc=6198262 |doi=10.12688/f1000research.14385.1 |doi-access=free }}</ref> Many organisms rely on their symbionts in order to develop properly, this is known as co-development. In cases of co-development the symbionts send signals to their host which determine developmental processes. Co-development is commonly seen in both arthropods and vertebrates.<ref name="Li-2015"/>
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