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===Limbs and locomotion=== Unlike ''P. robustus'', the forearms of ''P. boisei'' were heavily built, which might suggest habitual [[suspensory behaviour]] as in [[orangutan]]s and [[gibbon]]s.<ref name=McHenry2007/><ref name=Dominguez2013/><ref name=Lague2019/> A ''P. boisei'' [[shoulder blade]] indicates long [[infraspinatus muscle]]s, which is also associated with suspensory behavior.<ref>{{cite journal|first1=D. J.|last1=Green|first2=H.|last2=Chirchir|first3=E.|last3=Mbua|year=2018|title=Scapular anatomy of ''Paranthropus boisei'' from Ileret, Kenya|journal=Journal of Human Evolution|volume=125|pages=181β192|doi=10.1016/j.jhevol.2017.06.013|pmid=30502893|doi-access=free|bibcode=2018JHumE.125..181G }}</ref> A ''P. aethiopicus'' ulna, on the other hand, shows more similarities to ''Homo'' than ''P. boisei''.<ref name=Lague2019>{{cite journal|first1=M. R.|last1=Lague|first2=H.|last2=Chirchir|first3=D. J.|last3=Green|first4=E.|last4=Mbua|year=2019|title=Humeral anatomy of the KNM-ER 47000 upper limb skeleton from Ileret, Kenya: Implications for taxonomic identification|journal=Journal of Human Evolution|volume=126|pages=24β38|doi=10.1016/j.jhevol.2018.06.011|pmid=30583842|s2cid=58607106|url=https://www.researchgate.net/publication/329486548|doi-access=free|bibcode=2019JHumE.126...24L }}</ref> ''Paranthropus'' were [[biped]]s, and their hips, legs and feet resemble ''A. afarensis'' and modern humans.<ref>{{cite journal |vauthors=Wood B, Richmond BG |title=Human evolution: taxonomy and paleobiology |journal=Journal of Anatomy |volume=197 |issue=1 |pages=19β60 |date=July 2000 |pmid=10999270 |pmc=1468107 |doi=10.1046/j.1469-7580.2000.19710019.x}}</ref><ref>{{cite journal|first1=T. M.|last1=Ryan|first2=K. J.|last2=Carlson|first3=A. D.|last3=Gordon|display-authors=et al.|year=2018|title=Human-like hip joint loading in ''Australopithecus africanus'' and ''Paranthropus robustus''|journal=Journal of Human Anthropology|volume=121|pages=12β24|doi=10.1016/j.jhevol.2018.03.008|pmid=29706230|s2cid=14060188|url=https://www.repository.cam.ac.uk/handle/1810/277557|doi-access=free|bibcode=2018JHumE.121...12R }}</ref> The pelvis is similar to ''A. afarensis'', but the hip joints are smaller in ''P. robustus''. The physical similarity implies a similar walking gait.<ref>{{cite journal |vauthors=Macchiarelli R, Bondioli L, Galichon V, Tobias PV |title=Hip bone trabecular architecture shows uniquely distinctive locomotor behaviour in South African australopithecines |journal=Journal of Human Evolution |volume=36 |issue=2 |pages=211β32 |date=February 1999 |pmid=10068067 |doi=10.1006/jhev.1998.0267|bibcode=1999JHumE..36..211M }}</ref> Their modern-humanlike big toe indicates a modern-humanlike foot posture and range of motion, but the more distal ankle joint would have inhibited the modern human toe-off [[bipedal gait cycle|gait cycle]]. By 1.8 mya, ''Paranthropus'' and ''[[H. habilis]]'' may have achieved about the same grade of bipedality.<ref>{{cite journal|first1=R. L.|last1=Susman|first2=T. M.|last2=Brain|year=1988|title=New first metatarsal (SKX 5017) from Swartkrans and the gait of ''Paranthropus robustus''|journal=American Journal of Physical Anthropology|volume=77|issue=1|pages=7β15|doi=10.1002/ajpa.1330770103|pmid=3189526}}</ref>
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