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==Fossil record== The oldest [[fossil]] that can be related to Cornaceae is β ''Hironoia fusiformis'', an [[extinct]] taxon collected from the [[Futaba Group]] sediments at Kamitikaba, [[Japan]]. [[Synapomorphies]] of the fruits of the fossil occur also in extant Cornaceae. The age of the sediments is of early [[Coniacian]] to early [[Santonian]] (about 88 Ma).<ref>The Timetree of Life edited by S. Blair Hedges and Sudhir Kumar, OUP Oxford, 23. apr. 2009 - 576 pages, {{ISBN|0191560154}}, 9780191560156.</ref> Although the mosaic of characters in ''Hironoia'' precludes assignment to an extant genus, the fiber rather than sclereid composition of the fruit places it within the [[Nyssaceae]]-Mastixiaceae. Other possible Cornaceae from [[Cretaceous]] sediments include [[endocarp]]s resembling [[Cornus]] from the [[Santonian]]-[[Campanian]] mesofossil assemblage of [[Γ sen]]. In slightly younger [[Late Cretaceous]] sediments ([[Maastrichtian]]) four genera of [[fossil]] mastixioid fruits (''Beckettia'', ''Eomastixia'', ''Mastixicarpum'' and ''Mastixiopsis'') have been described from [[Germany]].<ref>{{cite journal|last1=Takahashi|first1=M.|last2=Crane|first2=P.R.|last3=Manchester|first3=S.R.|year=2002|title=''Hironoia fusiformis'' gen. et sp. nov.; a cornalean fruit from the Kamikitaba locality (Upper Cretaceous, Lower Coniacian) in northeastern Japan|journal=Journal of Plant Research|volume=115|issue=6|pages=463β473|doi=10.1007/s10265-002-0062-6}}</ref>
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