Editing Centriole (section)

==History==
The centrosome was discovered jointly by [[Walther Flemming]] in 1875<ref>Flemming, W. (1875). Studien uber die Entwicklungsgeschichte der Najaden. Sitzungsgeber. Akad. Wiss. Wien 71, 81–147</ref><ref name="Bloodgood-2009">Bloodgood RA. From central to rudimentary to primary: the history of an underappreciated organelle whose time has come. The primary cilium. Methods Cell Biol. 2009;94:3-52. doi: 10.1016/S0091-679X(08)94001-2. Epub 2009 Dec 23. PMID 20362083.</ref> and [[Edouard Van Beneden]] in 1876.<ref>Van Beneden, E. (1876). Contribution a l’histoire de la vesiculaire germinative et du premier noyau embryonnaire. Bull. Acad. R. Belg (2me series) 42, 35–97.</ref><ref name="Bloodgood-2009"/> [[Edouard Van Beneden]] made the first observation of [[centrosomes]] as composed of two orthogonal centrioles in 1883.<ref>{{Cite journal | year = 2002 | doi = 10.1007/s00109-002-0374-y| pmid =  12226736 | issue =  9 | pages = 545–548| volume = 80| title = JMM - Past and Present| last1 = Wunderlich| journal = Journal of Molecular Medicine| first1 = V.| doi-access = free}}</ref> [[Theodor Boveri]] introduced the term "centrosome" in 1888<ref>Boveri, T. (1888). Zellen-Studien II. Die Befruchtung und Teilung des Eies von Ascaris megalocephala.
Jena. Z. Naturwiss. 22, 685–882.</ref><ref name="Bloodgood-2009"/><ref>Boveri, T. ''Ueber das Verhalten der Centrosomen bei der Befruchtung des Seeigel-Eies nebst allgemeinen Bemerkungen über Centrosomen und Verwandtes''. Verh. d. Phys.-Med. Ges. zu Würzburg, N. F., Bd. XXIX, 1895. [https://archive.org/details/bub_gb_UYcPAQAAMAAJ link].</ref><ref>Boveri, T. (1901). ''Zellen-Studien: Uber die Natur der Centrosomen. IV''. Fischer, Jena. [https://archive.org/details/zellenstudienbe00bovegoog link].</ref> and the term "centriole" in 1895.<ref>Boveri, T. (1895). Ueber die Befruchtungs und Entwickelungsfahigkeit kernloser Seeigeleier und uber die Moglichkeit ihrer Bastardierung. Arch. Entwicklungsmech. Org. (Wilhelm Roux) 2, 394–443.</ref><ref name="Bloodgood-2009"/> The [[basal body]] was named by [[Theodor Wilhelm Engelmann]] in 1880.<ref>Engelmann, T. W. (1880). Zur Anatomie und Physiologie der Flimmerzellen. Pflugers Arch. 23, 505–535.</ref><ref name="Bloodgood-2009"/> The pattern of centriole duplication was first worked out independently by [[Étienne de Harven]] and [[Joseph G. Gall]] c. 1950.<ref>{{cite book | last = Wolfe | first = Stephen L. | author-link = Stephen L. Wolfe | title = Biology: the foundations | publisher = Wadsworth | year = 1977|edition=First | url =https://archive.org/details/biologyfoundatio00wolf| url-access = registration | isbn = 9780534004903 }}</ref><ref>{{Cite book|volume=106 |year=1987 |pages=227–293|doi=10.1016/S0074-7696(08)61714-3 |title=The Centrosome and Its Role in the Organization of Microtubules |first1=I. A. |last1=Vorobjev |first2=E. S. |last2=Nadezhdina |series=International Review of Cytology|isbn=978-0-12-364506-7 |pmid=3294718}}. See also de Harven's own recollections of this work: {{Cite journal|title=Early observations of centrioles and mitotic spindle fibers by transmission electron microscopy|first=Etienne|last=de Harven |journal=Biology of the Cell |year=1994 |volume=80 |pages=107–109 |doi=10.1111/j.1768-322X.1994.tb00916.x|pmid=8087058|issue=2–3|s2cid=84594630|df=dmy-all}}</ref>