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==Early life== Ancel Benjamin Keys was born in [[Colorado Springs, Colorado|Colorado Springs]] on January 26, 1904, the son of Benjamin Pious Keys (1883β1961) and Carolyn Emma Chaney (1885β1960), the sister of actor and director [[Lon Chaney]] (1883β1930).<ref name='Brody2004'>{{cite news | first = Jane E. | last = Brody | title = Ancel Keys, 100, Promoter of the Mediterranean Diet, Dies | date = November 23, 2004 |url=https://www.nytimes.com/2004/11/23/obituaries/23keys.html | work = [[The New York Times]] | access-date = February 5, 2011}}</ref> In 1906 they moved to San Francisco before the [[1906 San Francisco earthquake]] struck.<ref name='Sullivan2004'>{{cite news | first = Patricia | last = Sullivan | title = Ancel Keys, K Ration Creator, Dies | date = November 24, 2004 |url=https://www.washingtonpost.com/wp-dyn/articles/A7213-2004Nov23.html | newspaper = [[The Washington Post]] | access-date = February 5, 2011}}</ref> Shortly after the disaster, his family relocated to [[Berkeley, California|Berkeley]] where he grew up. Keys was intelligent as a boy; [[Lewis Terman]], a noted psychologist and inventor of the [[Stanford-Binet IQ Test]], described him as intellectually "gifted".<ref name ='Sullivan2004'/> During his youth, he left high school to pursue odd jobs, such as shoveling bat guano in Arizona, being a [[powder monkey]] in a Colorado mine, and working in a [[lumber camp]].<ref name='Hoffman1979'>{{cite web |url=http://mbbnet.umn.edu/hoff/hoff_ak.html | title = Meet Monsieur Cholesterol | access-date = February 5, 2011 | last = Hoffman | first = William | year = 1979 | work = [[University of Minnesota]] | publisher = University of Minnesota}}</ref> He eventually finished his secondary education and was admitted to the University of California at Berkeley in 1922.<ref name=Hoffman1979/> ===Higher education=== At the [[University of California, Berkeley]], Keys initially studied chemistry, but was dissatisfied and took some time off to work as an [[Oiler (occupation)|oiler]] aboard the [[American President Lines]] ship SS ''President Wilson'', which traveled to [[China]].<ref name=Hoffman1979/> He then returned to Berkeley, switched majors, and graduated with a B.A. in [[economics]] and [[political science]] (1925) and M.S. in [[zoology]] (1928).<ref name=Hoffman1979/> For a brief time, he took up a job as a management trainee at [[F. W. Woolworth Company|Woolworth's]], but returned to his studies at [[Scripps Institution of Oceanography]] in [[La Jolla, California|La Jolla]] on a [[scholarship|fellowship]]. In 1930, he received his Ph.D. in [[oceanography]] and [[biology]] from UC Berkeley.<ref name=Hoffman1979/> He was then awarded a [[United States National Research Council|National Research Council]] fellowship that took him to [[Copenhagen, Denmark]] to study under [[August Krogh]] at the Zoophysiological Laboratory for two years.<ref name=Hoffman1979/><ref name='Zadunaisky1969'>{{cite book | last1 = Zadunaisky | first1 = JA | title = Fish Physiology | chapter = The Chloride Cell: The Active Transport of Chloride and the Paracellular Pathways | volume = 10 | editor1= William Stewart Hoar |editor2=David J. Randall | publisher = Academic Press | year = 1969 | pages = 130β136 | chapter-url=https://books.google.com/books?id=8fuEB7O3IqIC&q=ancel%20keys%20willmer&pg=PA130 | access-date = February 5, 2011 | isbn = 978-0-12-350430-2 | quote = Ancel Keys was a fellow of the National Research Council of America, who did the perfusion experiment leading to the observation of chloride secretion in the Zoophysiological Laboratory in Copenhagen, Denmark, under the direction of August Krogh.}}</ref> During his studies with Krogh, he studied fish physiology and contributed numerous papers on the subject.<ref name=Zadunaisky1969/> Once his fellowship ended, he went to [[University of Cambridge|Cambridge]] but took some time off to teach at [[Harvard University]], after which he returned to Cambridge and earned a second Ph.D. in [[physiology]] (1936).<ref name ='Hoffman1979'/>
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